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Rozwój poglądów na przynależność systematyczną niektórych rodzajów rodziny Anomalinidae Cushman

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SUMMARY

The representatives of the family AnomaUntdae C u s h m a

n

1927 are characterized by both vast geographicai disU:ibution and wide stratlgraphic range. They become the subject of extensive studies since the XIX c. They were usually described under various generic names, and when Reuss (1862) pro-posed the family Rotaliidae, several ~ener~ of ~o­ chospirally coiled forms were placed m thIS famlly. The family AnomaZinidae was proposed by CUshman (1927) for the forms with aperture equatorial and sometimes extending on test walls. The latter fa-mily, as interpreted by CUshman, also comparised som£' highly specialized forms with modified' aper-tUre.

'The family Anomalinidae is genetically' related to the family Rotaliidae E h r e n b erg. According to Cushman (1948), the transitional link is represen-ted by the genus Discorbis, and particul~ly by its representatives with chitinous lining at early onto-'genic stages and traces of agg~utinated layer. HC?we-ver, some typical representatives of Anomalinidae, as e.g. the genus Gavelinella, were placed in the family RotaZiidae even by Cushman. Some ~ub­ 'sequent authors rejected the family AnomaUnidae

(see e.g. Galloway, 1933, Glae.;sner, 1945; Brotzen, 1942) whereas others accepted it with some modi-fications (e.g. Vassilenko, 1954, 1959; Loeblich and Tappen. 1964).

According to the present autor, the genera placed by Loeblich and Tappan (1964) in the subfamily AnomaZininae of the family AnomaUnidae should be separated in two subfamilies because of differences in location of the lateral part of the aperture. The forms with interiomarginal-equatorial aperture ex-tending on the ventral side, those with the aperture situated on ventral side, as well as those with aper-ture terminal but related vlith ventral side during the ontogeny, should be left in the subfamily Anomali-ninae. The forms with the aperture extending on the dorsal side, and their allies with a modified aperture, should be placed in a new subfamily. The rich assemblage of foraminifers from, the Upper Albian-Lower Turonian of the Polish Lowland was assigned to the family of Anomalinidae after' a de-tailed analysis of the position of embryonic chamber in the test, the mode of test coiling, degree of evo-lutness of 'the' test, location and cou):'se of lateral part of the aperture, and microstructure of test wall.

PE310ME

ITpe~cTaBHTeJlH ceMeJicTBa AnomaZintdae C u,s

h-m a n, 1927 XapaKTepHsYEOTCfi WHpOKHM BepTHKaJ1b-HhIM HHTepBaJIOM H 06l.UHpHOA DJlO~a;qblO pacnpoCT-paHeIiHfi. B CBfi3H C 9THK OHH npHBJleKaJIH BHHMaHHe HCCJIe,!J;oBaTeJleA em;e B npOWJlOM CTOJ1eTHH. 'YnoMfi-HYTbIe q,opaMHHHq,epbI OIIHCl.IB8J1H(''b JI<l,!J; pa3HhIMH PO,ll;OBbIMH Ha3BaHHHMH, a DOCJle c03,!J;anHH PeyccoM, (1862) ceMeAcTBa Rotaliidae K HeMY Ha,'IaJIH OTHOCHTb MHOrO'IHCJIeHHhIe pO,ll;hI TpoxocnHpaJlbHO cBepHYThIx cpoPM. CeMeAcTBo Anomalinidae 6hIJIO 06pn30BaW) KywMaHoM (1927) Ha OCHOBaH~ .poPM C 3KRaTOpHa-JlbHhIM DOJIOllteHHeM YC'l'bH, pacnpoCTpaHSBTUHMCH ID!Or,ll;a Ha CTeHKH paKoBHHhI. KpoMe Toro, K 9TOMY ceMeAcTBY OH 3a'IHCJIHJI BbICOKO pa3BHTbIe cP0PM1!I C MO,n:Hq,HD;HpOBaHHblM YCTbeM. CeMeAcTBO .1numaZinidae reEeTH'IeCKH CBH3aHO C ceMeAcTBoM Rotaliidae E

h-i:

e n b erg. Coe,lJ;HHHIO~HM 'IJIeHOM, KaR: YKa3bIBaeT KywMaH (1948), fiBJIHeTCH pO,ll; DiBCOTbi.~, :El oco6eH-HOCTH TaKHe npep;cTaBHTeJIH STOro pOAa, KOTopLle 06-JIa,ZJ;amT XHTHHOBOA nO,ll;KJIa,!J;KoA Ha paHHHX CTa,ll;KHX oHToreHeTH'IeCKOrO pa'3BHTJoJH H CJIe;n;aMH HaJIH<!HIl: ar-rJIIOTHHHPQBaHuoro CJIOH. CaM KYWKaH OTHOCHJI ewe K ceMeAcTBY Rotatiidae THnM'IHbIX Dpe,IICTaBHTeJ1eti1: ceMeAcTBa Anomalinidae (HanpHMep, P(\,!J; Govelinella). B ;n;an:bHe:il:weM O;!lHH HCCne110BaTeJlH ne I1pH3HaBaJIH

('p~eAc'l'Ba Anomalinidae (reJIJtoBeA, 1933; rJIeCCHep,

194!l: BpoTueH, 1942 H }Jp.), ,ll;pyrHe lit£> (BaccJolJleHKO,

1954, 1959; JIe6J.1HX H TanneR, 1964) npHHHMa.lIH ero, O,ll;HaxO B 'IieCKOJIbKO HHOM nOHHMaHHH.

ITo MHeHHEO aBTOpa, POAbI DOMe~eHHbIe B CHCTe-MaTHKe JUi6JIHXa H TanneHa (1964) B ceMeticTBe Ano-malinidae, nO,ll;ceMeAcTBe AnomaZininae Ha OCI)OSaHMH nOJI02KeHHfi 60KOBOH 'IaCTH ,YCTWI', ,ll;OJl1KIH,I 0'l'110CHTCH K ,ll;BYM nO,!J;ceMeHCTBaM. «POPMbI C 9KBaTOpHQJIbHbIM 6a3aJIbHhIM YCTbeM, nepexo,!J;fim;HM Ha 6pIOII~HYIO CTO-posy, .poPMbI. C YCTheM Ha 6PEOWHo:il: C'TOpOHe H .pop .• MDI C KOHe'IHbIM YCTbeM, HO OHTOreHeTH'IeCKl'.! CIIH3aH-HhIM C 6pmwIwA CTOPOHOti paKoBHHbI, CJIe,llYCT OCTa-BHTb B nO,ll;ceMetiCTBe AnomaIininae. ));.1JH <pOPM C yc-'IbeM nepeXO,!J;Hm;HM Ha CIIHHHym CTOpOHY JoL ;!lJIH PO;ll;-CTBeHHhIX HM q,OPM C MO,ll;Hq,HIJ;HpOBilHHhIM YCTbeM CJIe,ll;yeT 06pa30BaTb HOBOe nO,ll;ceMeAcTBO. ITPH onpe-,ll;eJleHHH npHHap;JIelltHOCTH K ceMeACTBY AnomaLinidae MHorO'rHCJleHHbIX ee npe;!lCTaBMTeJIe:il:, pacnpOC'I'paHeH-HhIX B HHTepBaJIe BepxHHx aJIb6 - IImKHroi TYPOH Ra TeppHTopHH ITOJIbCKo:il: HM3MeHHOCTH, ,:a;eTaJIbHO aJIH3HpOBaJIHCb TaKHe npH3HaKH KaK nOJt02KeHHe Ha-'raJIbHOA K8MepI>I, cnoco6 cBepHYTOC'I'H paxOBHHKH, CTeneHb ee 3BOJlmTI!OCTH, nOJIO>KeHHe 60KOBO:il: 'IaCTH VCTbfi H MHKpoCTPYKTYpa CTeHKH,

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