v. 51—1/2: 153—208 K ra k ó w 1981
Stanisław L i s z k a , Janina L i s z k o w a
REVISION OF J. GRZYBÓWSKI’S PAPER (1896)
„FORAMINIFERA OF THE RED CLAYS FROM WADOWICE”
(PI. I—VI)
Rewizja pracy J. Grzybowskiego (1896) „Otwornice czerwonych iłów z Wadowic”
(PI. I—VI)
S tan isław L i s z k a , Janina L i s z k o w a : R evision of J. G rzybow ski’s paper (1896)
“Formaiinifera of the Red C lays from W.adoiwice”. Ann. Soc. Geol. Polomiae, 51—
—1/2: 153— 208, 1981 Kraików.
A b s t r a c t : T he foram inifers (115 species) described by J. G rzybowski (1896) com e from red shales (marls) and from green sh ales and m arls from W adowice.
(Polish Carpathians). G rzybow ski defined the age of the m icrofauna as Oligocene.
R evision of the species id en tified by G rzybowski points to the C am panian age of the red shales (marls) and to Paleocen e age of the green shales and marls.
K e y w o r d s : Foram inifera, Campanian, Paleocene, Carpathians.
Stan isław Liszka: In stytu t G eologii i Surow ców M ineralnych, A kadem ia G órniczo- H utnicza im. St. Staszica, Al. M ickiew icza 30, 30-059 Kraków, Janina Liszkowa:
In stytu t Geologiczny, Oddział Karpacki, ul. Skrzatów 1, 31-560 K raków
M anuscript received: April, 1980 accepted: May, 1980
T r e ś ć : O tw ornice (115 gatunków) opisane przez J. G rzybowskiego (1896) po
chodzą z czerw onych łupków (margli) oraz zielon ych łupków i m argli z Wa
dowic. J. G rzybowski określił w ie k tej m ikrofauny jako oligoceński. R ew izja ga
tunków oznaczonych przez G rzybowskiego w ykazała, że czerwone łupki (margle) należą do kampanu, a zielone łupki i m argle reprezentują paleocen.
— 154 —
INTRODUCTION
J. Grzybowski described m any new species of Foram inifera from the C arpathian flysch. He was the first investigator to point out the value of th e use of m icrofauna for stratigraphic correlation.
In 1896 he published in Polish a pap er entitled „Foram inifera of the Red Clays from W adowice” (Rozprawy W ydziału M atem atyczno-przy
rodniczego Akadem ii Umiejętności w Krakowie, vol. XXX, pp. 261—398, P lates VIII—XI). The soummary of th is paper, w ithout the description of species, appeared early in Germ an as “Foram iniferen d er rothen Tone von W adowice” (Bull. Pol. Acad. Sci., 1895, Kraków, pp. 305— 315).
The m a terial described by J. Grzybow ski is derived from a pit, inac
cessible today, near Wadowice in the Polish w estern Carpathians. The pit in question pen etrated the following beds (J. Grzybowski 1896, p. 264):
“ 1—26 m. Thin-bedded sandstones altern atin g w ith sandy shales rich in muscovite, containing sphaerosiderites in the lowermost part.
26—64 m. G rey claystones w ith intercalated m arls, w ith lustrous sm ooth surface, and w ith partings of green clay w ith abun
d an t pyrite.
64—70 m. Red clays (recte marls) w ith intercalations of w hite sugary- -grained sandstone.
70—80 m. Black oil (“bitum inous”) shales.
80—84 m. M enilite shales, w here the drilling was stopped”.
These beds w ere assigned to th e Subsilesian nappe by M. Książ- kiewicz (1951).
U nfortunately, the m aterial left by J, Grzybowski is incomplete. Its bulk is in th e collections of the In stitu te of Geological Sciences of th e Jagiellonian U niversity in Cracow, w hereas a p a rt of the m aterial is housed in th e M useum of N atural Sciences in Lwów (the form er Dzie- duszycki Museum), w here it was catalogued b y S. J. P asternak and W. T. Lewicki (1963).
The m icrofauna in the Cracow collection is poorly preserved, the specimens are v ery small and in m ost cases the p artic u lar species are represented b y single specimens. In this situation exam ination in im m ersion oil w as adopted w ithout the possibility of sectioning, w hich considerably hindered th e revision.
J. Grzybowski’s specimens, which w ere stored in glass test-tu b es w ere tran sferred to microscope slides. The test-tubes, along w ith th e ir original labels, have been archived. The collection preserved in the In s titu te of Geological Sciences Jagellonian U niversity, was catalogued by Z. M artini in 1971. Its registration n um ber is 115 P.
Since the p it w ith J. Grzybowski’s m aterial was filled up and it was
impossible - to obtain topotypes from there, Professor M. Książkiewicz showed us an outcrop a t Woźniki (near Wadowice) w here, in his opinion, the red m arls are eq uivalent to those from Wadowice. The microfafunal assemblages from sam ples collected a t Woźniki were, in fact almost identical to those described by J. Grzybowski.
The outcrop at Woźniki is situated east of th e village at a side road leading to an abandoned q u a rry (see fig. 4 in Książkiewicz, 1966).
Because there is still a divergence in opinions concerning th e age of the red “clays” from Wadowice (Upper Cretaceous to Oligocene?), and since several micropalaeontologists refer to J. Grzybowski’s paper (1896), a revision of this w ork seemed to be necessary.
Basing his observations on the available publications (dealing m ainly w ith the Palaeogene) amd the com parative m aterials from Nikoltschitz and W aschberg presented to him for use b y A. Rzehak, J. Grzybowski defined the age of th e “red clays” as Oligocene. The position of the
“red clays” and green m arls in the Wadowice p it over the Menilite shales corroborated his hypothesis. However, J. Grzybowski adm itted (p. 271) th a t w hen com paring th e m icrofaunal assemblages, he did not take into consideration 55 new species described by him self and la te r micropalaeontological studies showed th a t a p a r t of them w ere index species of the Upper Cretaceous.
W. Żelechowski (1923) describing the foram iniferal fauna from a bo
rehole at Lgota near Wadowice, regarded m icrofauna of the red clays from Wadowice to be of Eocene age.
M. F. Glaessner (1937) m aintained th a t th e specimens described by J. Grzybowski should be p a rtly assigned to th e Palaeocene and th a t th e y w ere comparable w ith the Palaeocene m icrofauna of the Caucasus.
Basing on the foram iniferal assemblage of J. Grzybowski, H. H ilter- m ann (1943) suggested an U pper Cretaceous or Danian age for the red
“clays” from Wadowice.
In 1950 M. Książkiewicz accepted the view of th e ir U pper Cretaceous age and noticed lithological sim ilarity of the red “clays” (correctly speaking marls) from Wadowice (p. 316) to the m arls of the Woźniki series. He regarded th e red m arls from Wadowice, overlying the Meni
lite shales, to be an o v erth ru st tectonic cap.
Citing the opinions of C.A. W icher (1943) and H. H ilterm ann (1943), R. N oth (1951) also assigned the red m arls from Wadowice to the U pper Cretaceous.
F. Huss (1957) com pared the m icrofaunal assemblage from the red
“clays” w ith the m a rly p a rt of the red beds from Węglówka, defining th e ir age as Low er Senonian (Santonian).
E. W. M jatliuk (1950) regarded th e red “clays” from Wadowice as an equivalent of th e Popiele Beds, b u t in h e r papers of 1966 and 1970 she changed h e r view, suggesting a Senonian age.
— 156 —
. A. H illebrandt (1962) believed th a t the m icrofauna from Wadowice showed close sim ilarity to th a t from the Lizard Spring Form ation and to the Cretaceous m icrofauna of the Caucasus reported by M. Glaessner (1937) and H. H. Subbotina (1950, 1953).
Giving descriptions of individual species, W. Griin et al. (1964), W. Griin (1969) determ ined the age of the red “clays” from Wadowice as Palaeocene.
E. J. K rajew a and B. F. Ziernecki (1969) and Y. Kiesel (1970) re
garded the beds in question to be of U pper Eocene age.
S. Geroch (1960) concluded th a t J. G rzybow ski’s m aterial is a m ix
tu re of U pper Cretaceous and Palaeocene microfauna.
In the Wadowice m icrofauna it is possible to distinguish the spe
cimens derived from red m arls (coloured in red) from the specimens collected from m arls and green clays (white specimens). Although G rzy
bowski trea ted his m aterial as coeval, he gave hints w heth er the p a rti
cular forms were derived from red “clays” or from green clays w ith marls.
As a resu lt of our revision, it appears th a t the specimens from the red clays rep resent U pper Senonian (Campanian) m icrofauna while the foram inifera from green clays w ith m arls a re assigned to the Palaeocene (see table 1) according to th e ir know n vertical range. In this revision, for space economy, all synonym s are omitted, save th e synonym s used by J. Grzybowski w hich are p u t w ith inverted commas. On th e other hand, the authors who described th e given species are cited. If a speci
m en is missing, or its state of preservation made the revision impossible, the corresponding species is cited in the table only. Grzybowski’s des
criptions of new species if already given in the “Catalogue of Foram ini
fe ra ” of B. Ellis and A. Messina (1940) 1, are not cited. Translated quota
tions from G rzybow ski’s original paper are m arked by inverted commas.
Generic nam es are given according to the classification of A. R. Loeblich and H. T appan (1964). The authors would like to express their thanks to Prof. M. Książkiewicz for valuable inform ation on the geology of the Wadowice region and for critical review of this paper and to D r S. Ge
roch for helpfull discussion and consultation.
T a b l e 1 Liist of Fonaminifera in the coll. of J. Gmzybowski (1896)
N am e used by J. Grzybowski Nam e used in this revision Red
“clays"
f Green clays with
marls
1. 2. 3. 4.
N ubecularia tibia J. et P. R eophax cf. nodulosa Brady, pars H yperam m ina dilatata
G rzybowski +
1 In the present te x t the abbreviation “Cat. E-M” is used.
1 2 3 4 Keram osphaera irregularis Psam m osphaera irregularis
Grzyb. + radiolaria sp. +
Psam m osphaera fusca Psam m osphaera fusca Schultze
Schultze +
Saccam m ina sphaerica Nodosaria sp. apertural cham
Brady ber +
H yperam m ina dilatata Rzk. H yperam m ina dilatata G rzy
bowski +
H. vagans Brady H yperam m ina ?nodata Grzy
bow ski +
H. nodata n. sp. H. ?nodata G rzybowski +
Rhabdam m ina abyssorum Rhizam m ina grzybow skii n. sp.,
M. Sars pars H yperam m ina sub-
discretiform is M jatliuk + R. subdiscreta Rzk. Hyperam m ina subdiscretiform is
M jatliuk +
R. linearis Brady Rhabdam m ina cylindrica
G laessner +
R. annulata Rzk. Reopax cf. 9ubnodulosa
Grzybowski + +
R eophax ovulum n. sp. Hormosina ovulum (Grzybow
ski) +
R. lenticularis n. sp. Radiolaria sp. +
R. duplex n. sp. var. a R eophax duplex G rzybowski +
R. d u plex n. sp. var. f3 R eophax pilulifera Brady +
R. pilu lifera Brady R. pilulifera Brady + +
R. guttifera Brady var.
scalaria R. scalaria G rzybowski +
Haplophragm ium Wazaczi A m m obaculites w azaczi (Grzy
Rzek. bowski) +
H. turpe n. sp. H aplophragm oides turpe G rzy
bow ski +
H. (Reussina) quadrilobum ?T,rochammina bulloidiform is
n. sp. (Grzybowski) +
H aplophragm ium (Reussina) Trocham m ina bulloidiform is
bulloidiform e n. sp. var. a (Grzybowski) +
H. (Reussina) bulloidiform e Trocham m ina globigeriniform is
var. (3 (J. et P.) +
A m m odiscus involvens Rss. A m m odiscus angustus Friedberg +
A. angygyrus Rss. A. angustus Friedberg + +
A. polygyrus Rss. A. angustus Friedberg +
A. sp. (fig. 36) Glom ospira gordialis (J. et P.) +
A. charoides P. et J. Gl. charoides (J. et P.) +
A. gordialis J. et P. Gl. gordialis (J. et P.) +
A. Schoneanus Siddal P lectin a sp. +
A. fa lla x Rzk. Trocham m inoides sp. +
A. aberrans n. sp. Specim en m issing
A gatham m ina dubia n. sp. Rzehakina epigona (Rzehak) +
— 158 —
1 2 3 4
Trocham m ina subglobulosa Haplophragm oides subglobu- -
n. sp. losus (Grzybowski) +
Tr. interm edia Rzk. T rocham m inoides interm edius
(Grzybowski) + + ■
Tr. pauciloculata Brady Cystam ina pauciloculata (Brady) + Tr. Carpenteri n. sp. Specim en indeterm inable
Tr. Carpenteri n. sp. var. Specim en indeterm inable angustior n. var.
Tr. subcoronata Rzk. Trocham m inoides coronatus
(Brady) +
Tr. acervulata n. sp. Trocham m ina acervulata
G rzybowski +
C yclam m ina suborbicularis H aplophragm oides (Cribrosto-
Rzk. moides) suborbicularis (Grzy
bowski) + +
C. retrosepta n. sp. Specim en m issing C. setosa n. sp. Specim en m issing , C. globulosa n. sp. Specim en m issing
T extu laria attenuata Rss. Spiroplectam m ina anceps
(Reuss) +
T. subhaeringensis n. sp. S. subhaeringensis (Grzybow
var. a ski) ?macrospheric form +
T extularia subhaeringensis Spiroplectam m ina subhaerin
var. P gensis (Grzybowski) ?micro-
spheric form +
T. flab elliform is Giimb. A ragonia ouezzanensis (Rey) + Plecanium sublim e n. sp. P lectin a coniform is (Grzybow
ski) +
T extu laria ca lix n. sp. Dorothia crassa (Marsson) + V erneulina abbreviata Rzk. Dorothia crassa (Marsson) + V. Szajnochae n. sp. R eussella szajnochae (Grzy
bowski) +
B igenerina nuda n. sp. Ellipsodim orphina subcom pacta
(Liebus) +
B. fa lla x Rzk. P lectin a len is (Grzybowski) +
Spiroplecta len is n. sp. P. lenis (Grzybowski) +
S. d eflex a n. sp. P. lenis (Grzybowski) +
Gaudryina pupoides d’Orb. G audryina m odica Berm udez + G. chilostom a Rss. Specim en indeterm inable
G. Schw ageri Rzk. Specim en indeterm inable C lavulina subparisiensis T ritaxia subparisiensis (Grzy
n. sp. bowski) +
A taxophragm ium conulus Specim en indeterm inable Rzk.
V irgulina d igitalis n. sp. P lectin a conversa (Grzybowski) -h
P leu rostom ella w adow icensis P leurostom ella w adow icensis
n. sp. G rzybowski +
P. Zuberi n. sp. Ellipsopolym orphina zuberi
(Grzybowski) +
1 2 3 4
P. sp. Specim en indeterm inable
Lagena globosa Walk. Specim en m issing L. subapiculata n. sp. Specim en m issing
L. orbignyana Seguenza Fissurina orbignyana Seguenza +
L. (Cidaria) cidarina n. sp. Fragm ent Nodosaria sp. + L. (Cidaria) coronata n. sp. Fragm ent Nodosaria sp. + G landulina subinflata n. sp. G landulina subinflata G rzy
bowski +
Nodosaria calomorpha Rss. N odosaria cylindracea Reuss + + Nodosaria pungens Rss. Specim en m issing
N. sim plicissim a n. sp. Fragm ents indeterm inable
N. cornuta Batsch N odosaria com atula Cushm an + N. alternans n. sp. Specim en indeterm inable
D entalina Boueana d ’Orb. Specim en m issing
D. subtilis Neugeb. D en talin a cf. com m unis (d’Orb.) + D. in d ifferen s Rss. D. cf. megalopolitania Reuss + D. laticollis n. sp. D. laticollis G rzybowski + D. acuticauda Rss. Ellipsonodosaria div. sp.
fragm ents +
D. d eflex a n. sp. E llipsoidella d eflex a (Grzy
bowski) +
D. verm iculum Rss. Specim en indeterm inable D. n. sp. indet. S pecim en indeterm inable
L ingulina dentata n. sp. L ingulina dentata G rzybow ski +
C ristellaria cym boides P yrulinoides cf. ellip tica Marie + + d ’Orb.
C. lunaria n. sp. Lagena? lunaria (Grzybowski) + C. concava n. sp. L enticulina concava (Grzy
bowski) +
C. Kochi Rss. L. cf. grata (Reuss) +
C. abscisa n. sp. L. abscisa (Grzybowski) +
R obulina kressenbergensis L. m acrodisca (Reuss) + Giimb.
R. subangulata Rss. var. L. subangulata (Reuss) + R. gracilis n. sp. Specim en m issing
R. cincta Grzyb. L enticulina cf. cultrata (Mont-
fort) +
R. pectinata n. sp. L. cf. cultrata (Montfort) + V aginulina n. sp. Specim en indeterm inable
F labellina n. sp. indet. N eoflabellina ju ven ile specim en + Polym orphina dubia n. sp. S pecim en indeterm inable
G lobigerina triloba Rss. G lobigerina triloculinoides Plum m er partim G. linaperta
(Finlay) +
G lobigerina bulloides d’Orb. Fragm ents G lobigerina sp.
partim G. varianta Subbotina and G. cf. velascoerkisis Cush
man +
Sphaeroidina austriaca Rss. Fragm ents Globigerina sp. +
— 160 —
1 2 3 4
P u llen ia com m unis d’Orb. Specim en indeterm inable
P. com pressiuscula var. Pu llen ia cf. jarvisi Cushm an + quadriloba Rss.
Truncatulina propinqua Rss. Cibicides richardsoni Berm udez + T. H antkeni Rzk. Specim en m issing
T. m irabilis n. sp. Specim en m issin g A nom alina com planata Rss. Specim en m issing
var.
A. parvula n. sp. Specim en m issing A. tenuis n. sp. Specim en m issing
Pu lvin ulin a Karreri Rzk. Eponides karreri (Grzybowski) +
P. H aidingeri d’Orb. E. cf. candidulus (Schwager) +
P. m egastom a Rzk. Gyroidina m egastom a (Grzybo
wski) +
P. subcandidula n. sp. Eponides subcandidulus (Grzy
bowski) + +
R otalia Rom eri Rss. C ibicides cf. proprius Brotzen + R. D unikow skii n. sp. N uttallides dunikow skii (Grzy
bowski) +
R. N iedzw iedzkii n. sp. C ibicidoides niedźw iedzki
(Grzybowski) +
* N odellum velascoense (Cush
man) +
* C harltonina florealis (White) +
* Stensioeina beccariiform is
(White) +
T he species m arked (*), not described by G rzybow ski him self, have been found in the w ashed m aterial included in to the G rzybow ski’s collection.
On the basis of the above list of species the age of red clays (marls) is Cam panian and of the green clays and m arls — Palaeocene.
Revision of the species preserved in the collection N u b e cu la ria tibia Jones et P ark er
Grzybowski J. 1896, p. 273, PI. VIII, Figs. 10— 11 (slide no 1); recte:
Reophax cf. nodulosa B rady and H yp e ra m m in a dilatata Grzyb.
PI. I, Figs. 5a, b
“Nub. tib ia Brady. Chall. rap. S. 135. Tab. I. Fig. 1—4"
“Length 0.8 mm. In th e red clay very scarse”.
In Grzybow ski’s collection th ree fragm ents are preserved, two of which correspond to the au th o r’s illustrations. According to the descrip
tion, these specimens have siliceous tests m ade up of qu artz grains. Due
to this fact they cannot be classified as belonging in the genus N u b e cularia, despite certain general resem blance of the shape of fragm ents to B rad y ’s illustrations (1884) of the species N u b e cu la ria tib ia Jones e t P a rk e r (Plate I, figs. 1—4).
Two of these fragm ents can be assigned to the species R e oph a x cf.
nodulosa B rady w h e re a s’the th ird specimen found in this slide belongs in the species H yp e ra m m in a dilatata (Grzybowski) (1896, P late VIII, fig. 17a, b).
W. Friedberg (1901) and M. Dyl^zanka (1923) cite after Grzybowski the species N u b e cu la ria tibia Jones et P a rk e r from the U pper C reta
ceous, yet th ey provide no illustrations.
Keram osphaera irre g u la ris Grzybowski (Cat. E. M.)
Grzybowski J. 1896, pp. 273, 274, Pl. VIII, Figs. 12— 13 (slide no 2, 2b, 3);
recte: Radiolaria sp. and Psammosphaera irre g u la ris (Grzybowski)
In Grzybowski’s collection, vials 2, 2b m arked "K eram osphaera ir r e g u la ris ” constain spherical siliceous forms w ith fragm ents of Radiolaria skeletons on the exterior.
In vial 3, u n d er th e nam e of K eram osphaera sp., specimens of the genus Psammosphaera have been found. They are spherical, w ith slight
ly rough exterior surface and no trace of ap ertu re. Test w all fairly thick, m ade up of fine quartz grains. W hite in colour. The specimens show no division into in te rn a l cham bers m entioned in G rzybowski’s description and illustration, W. Griin et al. (1964, p. 248) rig h tly uphold his species, changing only th e genus to Psammosphaera irre g u la ris
(Grzybowski).
Psammosphaera fusca Sehultze G rzybowski J., 1896, p. 274, PI. 8, fig. 14 (slide no. 4) ; recte: Psammosphaera fusca Sehultze.
PI. I. fig. 9
“Ps. fusca, Chall. rap. S. 249. Tab. XVIII. Fig. 1—8."
“T est spherical, m ade up of rela tiv ely coarse quartz grains, w h ich gives it a rough and b ristle aspect. A perture indistinct. D iam eter 0.4 mm. V ery rare in red clays.”
The specimens in J. G rzybowski’s collection are in accord w ith his description and illustration. They are included in the synonym s of Ps.
fusca Sehultze by: E. Luczkowska (1955), T. Neagu (1962,1970), U. P flau- m ann (1964), F. Huss (1966), W. Griin et al. (1966), W. Grim (1969), E. J. K rajew a, B. F. Ziernieckij (1969).
11 — R o czn ik PT G 51/1—2
— 162 —
Saccammina sphaerica B rady
Grzybowski J., 1896, p. 274, PI. VIII, fig. 155 (slide no. 5);
recte: Nodosaria sp.
PI. I. fig. 10a, b
“Sacc. sphaerica Brady. Chall. rap. S. 252. Tab. X V III. Fig. 11— 17. S a cca m m in a sphaerica. Rzehak. Foram. von N ikoltschitz. V erb. d. g. R. A. 1887. S. 87.”
A specimen found in J. G rzybowski’s collection and determ ined as S. sphaerica is the a p e rtu ra l cham ber of a specimen of the genus N o d o saria sp. A t high m agnification, radiate conical ap e rtu re is visible.
T. Neagu (1962) and F. Huss (1966) w ere therefore w rong to include this form in the synonyms of the species Saccamina sphaerica Brady.
H yp e ra m m in a dilatata Rzehak (Gat. E. M.) Grzybowski J., 1896, p. 274, PI. VIII, fig. 17a, b (slide no. 6);
recte: H yp e ra m m in a (Pelosina?) dilatata Grzybowski.
PI. I. fig. 8
The description and illustration in accord w ith th e specimens in th e collection. Since Rzehak gave no description and illustration of this form, according to th e rules of th e “In ternational Code of Zoological N om enclature”, J. Grzybowski is credited w ith th e authorship of this species as he was the first to give its description and illustration. This species was described from the Polish C arpathians by F. Huss (1966, p. 12) and H. Jurkiew icz (1967), y et th e ir specimens do not correspond to the holotype.
H yp e ra m m in a vagans B rady
Grzybowski J., 1896, p. 275, PI. VIII, fig. 18, (slide no. 7);
recte: H y p e ra m m in a ? nodata Grzybowski.
PI. I. fig. 7a, b
“H. vagan s Brady. Chall. rap. S. 260, T. X XIV . Fig. 1—9.”
In J. G rzybow ski’s collection th ere are a few tu b u la r fragm ents m ade up of fine sand grains. These are not attached forms, so th e y cannot be placed in th e species T o ly p a m m in a vagans (Brady) Thalm ann 1932— 3, b u t sooner in the species H yperam m ina? nodata Grzybowski.
A fter Grzybowski this species is cited u n d er the nam e of H. vagans by W. F riedberg (1902) and M. Dylqzanka (1923) who give v ery brief descriptions w ithout illustrations.
H yp e ra m m in a nodata n. sp. (Cat. E. M.) Grzybowski J., 1896, p. 275, VIII, fig. 16 (slide no. 8);
recte: H yp era m m in a? nodata Grzybowski, PI. I. fig. 6
“Quite sim ilar to the preceding form, but w ith distinct segm entation at regular intervals. R are in red clay.”
In the collection th e re are tw o fragm ents of tu b u la r form s w ith constrictions and rough surface, m ade up of fine qu artz grains. Dia
m e te r of fragm ents 0.2 mm. The author w ell illustrates one of them (fig. 16), b u t he does not bring out into relief its rough surface. V. Po- korny (1949) identified the species H . nodata w ith H. vagans, stating th a t in his m aterial from Nikolcice he found transitional form s betw een the tw o species and no traces of attachm ent. The species H . nodata Grzybowski was also reported by W. F riedberg (1901), M. Dylqzanka (1923), T. Homola, E. Hanzlikova (1955) M ariana Iva et al. (1971), O. Sa
m uel (1977). Because of th e sm all am ount of fragm ents preserved in the collection and th e ir small size, the generic determ ination is not certain.
R habdam m ina abyssorum M. Sars
Grzybowski J., 1896, p. 275, PL VIII, fig. 1—4 (slide no. 9);
recte: R h izam m ina g rz y b o w s k ii n. sp. pars H yp e ra m m in a subdiscreta (Grzybowski).
“Rhabd. a b ysso ru m . Brady. Chall. rap. S. 266. Tab. X X I.”
“Found only as fragm ents o f tests. Tube m ade up of fin e sand grains w ith rough or b ristle surface, straight or bent, w ith ou t constrictions. D iam eter 0.2— 0.6 mm. L ength of fragm ents 0.8— 1.2 mm, rarely up to 2 mm. Fig. 3 cor
responds to Rh. a b ysso ru m var. irregularis Carp, described by Brady. In R zehak’s m aterial from N ikoltschitz partly as Rh. emaciata. Common in red
clay.”
In the collection there are few tu b u la r, non-flattened form s w hich, considering the test stru ctu re, can be assigned to tw o species: one re p re sented by forms w ith a not v ery rough surface, for w hich a new specific nam e is suggested (described below); the other represented by a few fragm ents of tu b u la r forms averaging 1.00 m m in length and 0.25—
0.30 m m in thickness, w ith v e ry rough surface, walls m ade up of coarse, angular quartz grains cemented w ith a sm all am ount of ferrous-siliceous m aterial. These fragm ents are identical w ith those found in a phial m arked Rhabdam m ina subdiscreta Rzehak (slide no. 10). The la tte r species of G rzybowski E. M jatliuk (1970) includes in the synonym s of the new ly determ ined species H yp e ra m m in a subdiscretiform is. The p re sent authors are of th e opinion th a t according to th e rules of the “In te r
national Code of Zoological N om enclature”, th e nam e H yp e ra m m in a sub
discreta (Grzybowski) should be preserved.
ii*
— 164 —
Several au th o rs (W. Friedberg 1902, M. D ylążanka 1923, R. Noth 195-1, A. H illebrandt 1962, H. Jurkiew icz 1967) assign J. G rzybowski’s species (1896) to the synonym s of Rh. abyssorum M. Sars, and W. Griin et al. (1964) place it w rongly in Psam m osiphonella rze haki Andreae.
T. Neagu (1962) and W. G riin et al. (1964) include Rh. abyssorum var.
irre g u la ris (non Carpenter) Grzybowski .(PI. VIII, fig. 3) in the synonyms of R h isa vn n in a in d iv isa Brady.
K hizam m ina g rz y b o w s k ii n. sp.
1896 R habdam m ina abyssorum M. Sars, Grzybowski J., p. 275, Pl. VIII, fig. 1; Rh. abyssorum var. irre g u la ris Carp., fig. 3.
H o l o t y p u s : specimen presented in pl. I. fig. 1 a, b.
S t r a t u m t y p i c u m : red m arls (Campanian)
L o c u s t y p i c u s : Wadowice (J. G rzybow ski’s collection, 1896, slide no. 9a).
M a t e r i a l : 7 fragm ents (holotype and paratypes) from Wadowice, 20 fragm ents from Woźniki.
D i m e n s i o n s : length of fragm ents 1.35— 1.45 mm; diam eter 0.33—
0.45 mm; w all thickness 0.12— 0.15 m m
Tests tu b u lar, not flattened, w ithout constrictions, sometimes branched. W alls fairly thick, made up of v ery fine m aterial w ith sparse larg er grains of quartz, glauconite or pyrite. The test dissolves (com
pletely or p artly ) in hydrochloric acid. Observations in polarized light reveal the v ery fine-grained stru c tu re of walls composed of a base of carbonates w ith embedded quartz, glauconite or pyrite grains. In tern al channel refilled w ith calcite crystals. Test surface not very rough, m at, w ith p ro tru d in g sparse grains of the m entioned m inerals. Test light-grey to w hite in colour.
Among th e preserved specimens, form s shown by J. Grzybowski in figs. 1 and 3 have been found. Specimens branching dichotomously, like those in fig. 3, are also common in the m aterial from Woźniki. N either J. Grzybow ski’s m aterial nor th a t from Woźniki contains specimens w ith a spherical central chamber.
R habdam m ina subdiscreta Rzehak (Cat. E. M.) Grzybowski J., 1896, p. 275. Pl. VIII, figs. 5—6 (slide no. 10);
recte: H yp e ra m m in a subdiscreta (Grzybowski).
Pl. I. fig. 2 a, b
"Rh. subdiscreta. Rzehak. Foram iniferen von N ikoltschitz. Verh. d. g. R. A.
1887. S. 87.”
In the collection th ere are 3 fragm ents of tu b u la r form s open a t both ends. Walls composed of angular qu artz grains of varying size, w ith
little siliceous cem ent. Surface v ery rough. Two of these specimens are well illustrated by the author, w ith m arked constrictions. In transm itted light also narrow ings of the channel are visible, w hich do not form sutures.
This species was introduced a fte r J. G rzybow ski (1896) as “sub
d iscreta” by W. F riedberg (1901) and M. D ylążanka (1923). E. W. M jat- liuk (1950) w rongly assigned the species R h. abyssorum M. Sars. p re
sented by J. Grzybowski (1896) on pl. VIII, fig. 4 and R h subdiscreta Rzehak on pl. VIII, figs. 5—6 to the new species H yp e ra m m in a g v id o - nensis n. sp. In h er paper from 1970, this au th o r places Я. subdiscreta Rzehak in the new species H yp e ra m m in a subdiscretiform is M jatliuk.
Rhabdam m ina linearis B rady
Grzybowski J., 1896, pp. 275—276, Pl. VIII, fig. 7; (slide no. 11).
recte: Rhabdam m ina c y lin d ric a Glaessner Pl. I. fig. 3
“Rh. linearis Brady. Chall. rap. S. 269. T. X X L Fig. 1—4” (lapsus) Pl. X X I figs. 1—6).
“Fragm ent of a thin, narrow tubule, straight, not w id en in g into a chamber, with rough surface, w ithout constrictions. D iam eter 0.2 mm. Rare in m arls. ”
In the collection there is one tu b u la r specimen w ith the rough su r
face and walls composed of angular grains of hyaline, translucent quartz. Internal d iam eter of the channel v ery small. There are no thick
enings or narrow ings on the whole length of the fragm ent, which fact was stressed and illustrated by the author. The specimen is slightly tran sp aren t, which was already stressed by W. F riedberg (1902) and m entioned by A. G riin et al. (1964) in his description of the species Psamm osiphonella c y lin d ric a (Glaessner), in which he includes as a syno
nym the form described by J. Grzybowski (1896) u nder the nam e “ Rh.
linearis B rady”. M. J. Glaessner (1937) regards Rh. linearis (non Brady) Grzybowski (1896) as a synonym of his species Rh. cylindrica, giving a com prehensive description corresponding to G rzybowski’s specimen.
R. Noth (1951) supposes erroneously th a t the species of Grzybowski and Glaessner can belong in Rh. abyssorum . H. Jurkiew icz (1967) assigned to the species Rh. linearis B rady both the fragm ents w ith characteristic thickenings, corresponding to B rady’s description and illu
strations (1884), and the specimens w ithout these thickenings placed in this species by J. Grzybowski (1896) as he thought th a t these fragm ents are presum ably th e segments betw een the thickenings. S im ilarly G rzy
bowski (1897, p. 275) points out th a t specimens assigned by him to Rh. linearis B rady show slight inflations in th e centre th a t form an oval cham ber. It is w o rth noting, however, th a t in the assemblage from th e Carpathians,, form s identical w ith Grzybowski’s specimen (1896) i.e. up
— 166 —
to 0.2 mm in diam eter, coarse-grained w ith little cement, hyaline, slight
ly translucent w ithout thickenings, usually occur in th e Palaeogene and are reported u n d e r th e specific nam e of R h. c y lin d ric a Brady. N. J. Ma- slakowa (1955), A. H illebrandt (1962) and E. J. K rajew a, B. F. Zarneckij
(1969) are of th e same opinion, including Grzybowski’s form in the syno
nyms. Also E. W. M jatliuk (1970) regards Grzybow ski’s specimen as a synonym of H yp e ra m m in a c y lin d ric a (Glaessner). The assignation of this species to the genus H yp e ra m m in a seems to be incorrect.
Rhabdam m ina annulata Rzehak (Cat. E. M.) Grzybowski J., 1896, p. 276, PI. VIII, fig. 8, 9 (slide no. 12);
recte: R eophax cf. subnodulosa Grzyibowski.
PI. I. fig. 4 a, b, c
The collection contains two fragm ents corresponding to the speci
m ens illu strate d by th e author. E xam ination in tran sm itte d light re vealed th a t th e y both belong in the genus R eophax. One of them , 0.9 mm in len g th is composed of 6 cham bers in a straig h t linear series (fig. 9). The em bryonal cham ber is spherical, 0.175 m m in diam eter, la te r cham bers are pyriform and connected by thin pipes.
The ap e rtu ra l cham ber, 0.25 m m in diam eter, has a term inal ap ertu re.
The other specim en (fig. 8) is sm aller, slightly curved, composed of 3 cham bers w ith o u t th e em bryonal and ap e rtu ra l cham bers. Length of this fragm ent 0.6 mm, thickness 0.2 mm. The surface rough, walls of fine quartz grains w ith siliceous cem ent. L ight-grey colour of th e fragm ent suggests th a t th e two specimens come from green m arls. In his description of the species "R habdam m ina annulata Grzybowski”, T. Neagu (1962) suggests th a t the species described and illustrated by J. Grzybowski (1896) should be placed in th e genus Reophax. The spe
cim en exam ined bears closest resem blance to the species R e ophax s u b - nodulosa G rzybow ski (1897). In consequence, the inclusion of the speci
m ens described and illu strated by Grzybowski in th e synonym y of Rh.
annulata Grzybowski (T. Homola, E. Hanzlikova 1955, H. Jurkiew icz 1967, O. Sam uel 1977) or in the species Psam m osiphonella annulata (Andreae) (W. Griin et al. 1964) does not come into consideration.
R e ophax o v u lu m n. sp. (Cat. E. M.)
Grzybowski J., 1896, p. 276, PI. VIII, figs. 19—21 (neotyp slide no. 13);
recte: H orm osina o v u lu m (Grzybowski).
No specimens of this species are preserved in the collectien, only one specimen has been found in the rem aining, non-determ ined m aterial (vial no. 67 w ith Lagena cidarina). The species in question, v ery common in the Cretaceous and Palaeocene of the C arpathians, was described and cited under th e nam e of H orm osina o v u lu m (Grzybowski) by M. Glaess-
n e r (1937), R. Noth (1951), V. VaStfek (1951), S. Geroch (1955, 1959, 1960, 1966), T. Homola, E. Hanzlikova (1955), N. J. Maslakowa (1955), J. Mor- giel (1959), A. H illebrandt (1962), W. G riin e t al. (1964), U. P flaum ann (1964), E. Hanzlikova (1965), H. Jurkiew icz (1967), T. Neagu (1979), J. Sandulescu (1972), O. Sam uel (1977). E. W. M jatliuk (1966, 1970) placed this species in th e new ly preposed genus C arp ath iella . J. Szcze-
chura, K. Pozaryska (1967) follow th e same way.
R e o ph a x le n ticu la ris n. sp. (Cat. E. M.) G rzybowski J., 1896, p. 276, PI. VIII, fig. 22 (slide no. 14);
recte: R adiolaria sp.
F airly num erous specim ens determ ined in the collection as R eophax le n ticu la ris rep resen t len ticu lar Radiolaria, fairly abundant in th e T er
tia ry Flysch form ations. This fact was already m entioned by S. Geroch, R. G radzinski (1955), S. Geroch (1960), who gave an accurate description and illustrations.
R eophax d u p le x n. sp., var. alfa (Cat. E. M.)
Grzybowski J., 1896, p. 276, PI. VIII, figs. 23, 24 (slide no. 15);
recte: R eophax d u p le x Grzybowski.
PI. I, fig. 12
Two specimens in th e collection conform closely w ith the a u th o r’s description and p a rtly w ith his illustration, b u t only one of them is presented accurately in fig. 23. Its dimensions are: length 1.1 mm, dia
m e ter 0.75 mm. The o th er specimen does not correspond to fig. 24. It is m uch sm aller, cham bers spherical of n ea rly equal size, distinctly sepa
rated. Its dimensions are: length 0.6 mm, d iam eter 0.45 m m . Walls of both specimens are m ade up of coarse, angular qu artz grains, the surface very rough. A pertures invisible. In spite of this, the two speci
mens can be included in the species R e ophax d u p le x Grzybowski. The specimen illustrated b y J. Sandulescu (1972, fig. 7) can also be assigned to this species. It does n ot seem rig h t to identify this species w ith R. p ilu life r B rady (F. Huss 1966, A. Jednorow ska 1968).
R e ophax d u p le x n. sp. var. beta (Cat. E. M.) Grzybowski J., 1896, p. 277, PI. VIII, fig. 25 (slide no. 16);
recte: R eophax p ilu life r Brady.
PI. I, fig. 13
In J. G rzybowski’s collection th e preserved specim en corresponds to the description and figure of the author. I t is composed of 2 cham bers of unequal size m ade up of sand grains, walls not so rough as in R. d u p le x.
— 168 —
In view of th e different character of the surface and the size of cham bers, it should be assigned to R . p ilu lije r a Brady. Placing of this v arie ty in the species R. d u p le x, as suggested by Grim et al. (1964), E. W. M jat
liuk (1970) and T. Neagu (1970), is incorrect.
R eophax p ilu lije r a Brady
Grzybowski J., 1896, p. 277, Pl. VIII, figs. 27—28 (slide no. 18);
recte: R e ophax p ilu life ra Brady.
Pl. I, fig. 14
41 Rh, pilulifera Brady I.e. S. 291. Tab. X X X . Fig. 12— 17. Rh. pilulifera Rzk.
Verh. d. g. R. A. 1887. S. 87. 1888. S. 191, 192.”
“Test of three spherical chambers, arranged like in D entalina, m ade up of fine quartz grains, w ith bristle surface. The cham bers enlarge rapidly, th e last one som etim es occupies half of the test. A perture in the form of a circular opening at the top of th e lastform ed chamber. Length 0.5— 1.5 mm. V ery scarce in m arls and red clay.”
In the collection th e re is only one specimen corresponding to th e a u th o r’s description and figure 27. Length of the whole specimen com
posed of 3 cham bers is 1.25 mm, d iam eter of th e initial cham ber 0,35 mm, of the m iddle one 0.6 m m , th a t of th e a p e rtu ra l cham ber 0.7 mm. This species is v ery freq u en tly described in Polish and foreign m icropaleonto- logical publication. The form s described by J. Grzybowski from W ado
wice w ere included in the synonym s of this species by R. Noth (1912, 1951), W. G riin et al. (1964), U. P flaum ann (1964), F. Huss (1966), H. J u r kiewicz (1967). E. M jatliuk (1970) assigned them w rongly to the new species R. p a ra d u p le x n. sp.
R eophax g u ttife ra Brady, var. scalaria n. var. (Cat. E. M.) Grzybowski J., 1896, p. 277, pl, VIII, fig. 26 (slide no. 17);
recte: R eophax scalaria Grzybowski.
Pl. I, fig. 15
“Rh. gu ttifera. Brady. I.e. S. 295. Tab. X X X I. Fig. 10— 15.”
One preserved specimen corresponds to th e a u th o r’s description and figure. A fter Grzybowski this variety is reported by W. F riedberg (1901), M. Dylążanka (1923) and described by H. Jurkiew icz (1967). J. S chubert questions th e specific nam e of this v arie ty (1902, p. 21), m aintaining th a t such shape can be due to deform ation of the specimens. A. Griin et al. (1964) include G rzybowski’s v ariety in the synonym s of R. s p le n d - idus (Grzybowski) (1897, Pl. X, fig. 16), citing it from the paper of Friedberg (1901). It is w orth noting th a t figure 15 in plate 3 of A. Griin et al. (1964) conforms closely w ith J. G rzybowski’s specimen (1896).
Since, how ever, the nam e “splendidus” was introduced in the la te r
paper, the earlier nam e R. scalaria Grzybowski is retained. Also E. H anz
likova (1966) and O. Sam uel (1977) cite G rzybow ski’s species u n d er this name.
H a p lo p h ra g m iu m W azaczi Rzhk. (Cat. E. M.) G rzybowski J., 1896, 277, PI. VIII, fig. 29 (slide no. 20);
recte: A m m obaculites wazaczi (Grzybowski).
PI. I, fig. 11
The description corresponds to the figure and the preserved speci
men. Dimensions: length 1.5 mm, b read th of the ap e rtu ra l cham ber 0.6 mm, diam eter of the coiled portion 0.8 mm. Walls of very coarse angular light qu artz grains, th e surface of walls v ery rough. This spe
cies is sim ilar to A . c o p ro lith ifo rm is (Schwager) y et the la tte r species has more distinct sutures betw een more num erous cham bers, and the surface is not so rough. Since Rzehak gave no description or figure, J. Grzybowski is credited w ith th e authorship of this species, which is new ly described and illustrated by E. Hanzlikova (1972).
H a p lo p h ra g m iu m turpe n. sp. (Cat. E. M.)
Grzybowski J., 1896, pp. 277—278, PI. VIII, fig. 30 (slide no. 19);
recte: H aplophragm oides tu rp e (Grzybowski).
PI. I, fig. 16
One of the w ell preserved specimens corresponds to J. G rzybowski’s description and figure. Test small, involute, 0.6 m m in diam eter. Cham bers v ery indistinct, 6 in num ber, separated by slightly depressed su tures, slowly increasing in size. Slit-like ap e rtu re at the base of the last-form ed cham ber. Walls of coarse, sharp-edged grains of m ilky q u artz w ith little cem ent. The surface of walls rough. Except for J. G rzy
bowski (1897, 1901) and W. Friedberg (1901), not reported in the lite ra ture.
H a plo p h ra gm iu m (Reussina) q u a d rilo b um n. sp. (Cat. E. M.) Grzybowski J., 1896, p. 278, PL VIII, fig. 31 (slide no. 21);
recte: IT ro c h a m m in a b u llo id ifo rm is (Grzybowski).
PL I, fig. 17
T here is one specimen in the collection composed of three cham bers of nearly equal size, arranged in one plane. Surface very rough, walls of large, sharp-edged quartz grains. A p ertu re invisible. The longest dia
— 170 —
m e te r of th e 3-cham ber specimen 0.9 mm. The preserved specimen lacks the fo u rth cham ber shown in fig. 31. I t is w orth noting th a t foram ini- fera corresponding to J. Grzybowski’s description and figure occur in th e m icrofaunal assemblage from the red shales from Woźniki alongside of specimens composed of th ree or m ore chambers. F. Huss (1966) des
cribes and figures sim ilar forms from W ęglówka (Santonian-M aastricht- ian), determ ining them as T ro ch am m in a b u llo id ifo rm is (Grzybowski).
Describing th e species Tro ch am m in a quadriloba (Grzybowski), S. Ge- roch (1960) includes Grzybowski’s H. q u a d rilo b u m in its synonyms, po
inting out th a t “the cham bers are trochospiral, increasing rapidly in size”, which observation is unconform able w ith the preserved specimens and Grzybow ski’s figure, as w ell as w ith specimens from Woźniki.
H a p lo p h ra g m iu m (Reussina) b u llo id ifo rm e n. sp. var. alfa (Cat. E. M.) Grzybowski J., 1896, p. 278, Pl. VIII, fig. 32 (slide no. 22 alfa);
recte: Tro ch a m m in a b u llo id ifo rm is (Grzybowski) Pl. I, fig. 16 a, b
In J. G rzybow ski’s m aterials th e re are four specimens composed of 3, 4 and 5 chambers. Cham bers globular, n early equal in size, 3-cham ber specimens have cham bers in a single plane, in 4-cham ber ones the fo u rth lies on 3 cham bers (fig. 31), w hereas in 5-cham ber specimens th e fifth cham ber lies on 4 chambers, w hich are in one plane (fig. 32). Walls fairly thick m ade up of large, angular quartz grains w ith little siliceous cement. The surface of walls v ery rough. N either in G rzy
bow ski’s specimens nor in the ones from Woźniki are any apertures to be seen.
Considering insignificant differences in the arrangem ent, F. Huss (1966) rig h tly includes both H. quadrilobum and H . b u llo id ifo re vai;.
alpha in the species Troch am m in a b u llo id ifo rm is (Grzybowski). Con
tr a r y to S. G eroch’s description (1960, p. 64) of T . quadriloba (Grzy
bowski), the specimens have never been found to be composed of m ore th a n 5 cham bers. Because of the sm all n u m b er of specimens in the collection, a polished section was m ade from a 3-cham ber specimen from Woźniki, which revealed the presence of links betw een the cham bers corresponding to the description and figure of F. Huss (1966).
H a p lo p h ra g m iu m q u a d rilo b um and H , b u llo id ifo rm e of J. Grzybowski w ere reported u n d er different generic nam es b y W. F riedberg (1901), R. Noth (1912), M. D ylążanka (1923), V. Pokorny (1949, 1953), V. Ho- mola, E. Hanzlikova (1955), F. Huss (1966), H. Jurkiew icz (1967) both from the Cretaceous and the Palaeogene of the C arpathian Flysch.
T. Neagu (1970) leaves the species Tro ch am m in a quadriloba and Tr.
b u llo id ifo rm e as valid, incorrectly including in the la tte r Tr. bulloidi- form e var. beta.
H a p lo p h ra gm iu m (Reussina) bu llo id ifo rm e var. beta n. sp. var.
Grzybowski J., 1896, p. 278, PI. VIII, fig. 33 (slide no. 22 beta) recte:
Tro c h a m m in a g lo b ig e rin ifo rm is (P ark er et Jones).
PI. II, fig. 1.
There is one specimen in th e collection belonging to th e genus T r o cham m ina. It is poorly preserved, p artic u larly th e last-form ed cham ber, b u t its stru c tu re corresponds to J. G rzybow ski’s description and figure.
Test composed of fine m aterial, surface less rough th a n in the previous species, w ith m ore cement. Cham bers trochospiral, rapidly increasing in size as in Tro ch am m in a g lo b ig e rin ifo rm is (P a rk e r et Jones), which was m entioned by J. Grzybowski.
Am m odiscus in vo lve n s Keuss
Grzybowski J., 1896, p. 279, PI. VIII, fig. 38 (slide no. 23); recte: A m m o- discus angustus (Friedberg)
PI. II, fig. 5.
“Operculina in v o lv e n s Rss. D enkschrift. d.k. Akad. d. W iss. in W ien, T. I.
S. 370. Tab. 46. Fig. 30.
C ornuspira in v o lv e n s R ss. Sitzungsber. d.k. Akad. in W ien. T. 48. S. 39. Tab. I.
Fig. 2.
Cornuspira in v o lv e n s Hantken. D ie Foram iniferen der C lavulina Szaboi S ch i- chten. S. 19. Tab. II. Fig. 2.”
“S iliceou s test of 8 planispiral coils corresponds in shape and arrangem ent to th e description and figu res o f Reuss. T h e coils are partly covered, the last one is the largest on e and m akes up nearly 1/4 of the diam eter of the test.
The m argin broad, n arrow ly rounded tow ard th e periphery, the sides concave.
D iam eter 0.8 mm. Scarce in red clays.”
Am m odiscus a n g y g y ru s Reuss
G rzybowski J., 1896, p. 280, PI. VIII, fig. 34 (slide no. 24 ab); recte:
Am m odiscus angustus (Friedberg)
PI. II, fig. 4.
“Cornuspira a n g y g y r a Reuss. D. s. d. A. W. Bd. I. S. 370. Tab. 46. Fig. 19.
C. a n g yg yr a Rss. S. b. d. A. W. Bd. 48. S. 38.”
“A lso this form does not d iffer in any respect from the one described b y R euss from th e M iocene and Septaria clays; it also has num erous low , flatten ed coils and th e surface only sligh tly rough. Scarce in m arls and red clays.”
— 172 —
Am m odiscus p o ly g y ru s Rss.
G rzybowski J., 1896, s. 280, tab. VIII, fig. 37 (slide no. 25); recte: Л т ш о - discus angustus (Friedberg)
Pl. II, fig. 3.
“Cornuspira p o ly g y ra Reuss. S. b. d. A. W. Bd. 48. S. 39. Tab. I. Fig. 2. C. p o ly - дуга H antken I.e. S. 19. Tab. Fig. 11.”
“T est rela tiv ely thin. N um erous low coils, periphery rounded, slig h tly concave on both sides. Scarce in red clays.”
One specim en of A . involve ns,p one of A . p o ly g y ru s and two of A. angygyrus are preserved in th e collection. All these specimens are identical. They have the same d iam eter and an equal num ber of coils (about 8) increasing evenly in size and overlapping. P erip h ery rounded.
Tests round or eliptic, bilaterally concave. The surface of walls lustrous, smooth. W alls thin, of cryptocrystalline siliceous m aterial. Test dia
m eter 0.75— 1.00 mm.
The specimens, and particu larly A . in v o lve n s, do not show any significant differences in thickness of the last two coils, w hich fact was noted by J. Grzybowski. A . p o ly g y ru s does not differ from the other form s in sm aller thickness of the test. Elliptical shapes of some spe
cimens m ay be due to lateral deform ation, which also causes flattening of the coils, p artic u larly pronounced on the last one, and sim ultaneous enlargem ent of the central depression (A . a n g y g y ru s , slide no. 24a, b).
W. F rie d b e r g . (1901) was the first to couple these species and assign them to the species C o rn u sp ira incerta d'Orb. suggesting th a t C. angusta an n. sp. can be its v arie ty (p. 637). W. G riin et al. (1964) also unites th e species described by J. Grzybowski and W. Friedberg, placing them in A . siliceus (Terquem). The la tte r Jurassic species has a larg er num ber of coils (15 on the average) and walls m ade up of coarse-grained m a
terial (vide Cat. E. М.). The specific nam es A . in v o lve n s and A . a n g y
g y ru s used by J. Grzybowski (1896) refer to calcareous species and therefore, m u st be discarded and so m u st the specific name A . p o ly g y ru s (Reuss). This la tte r Oligocene species has a considerably larg er num ber of coils (13— 15), and this featu re has been observed n eith er in J. G rzy
bowski’s specimens nor in the ones from Wożniki.
In her com prehensive description of G rz y b o w s k ie lla angusta (Fried
berg), E. W. M jatliuk (1970) includes A . in v o lve n s and A . a n g y g yru s of J. Grzybowski in its synonyms, leaving out w rongly A. polygyrus.
Moreover, th e range she gives for this species (Upper Senonian-Palaeo- cene) is too short. Identical forms reported from the Polish C arpathian u nder d ifferen t names, most commonly as A . incertus (d’Orb.) and A . siliceus (Terquem) have stratigraphic ranges from the Low er C reta
ceous to the Palaeogene (S. Geroch 1966).
Since the description and figures of. E. W. M jatliuk (1970) corres
pond to the preserved specimens from Wadowice, we accept h er nam e as valid.
Am m odiscus sp.
Grzybowski J., 1896, p. 280, PI. VIII, fig. 36 (slide no. 27); recte: G lo m o - spira gordialis (Jones et Parker).
Two specimens are preserved in the collection. One of them , shown in fig. 36, is G lom ospira gordialis (J. et P.), according to J. Grzybowski’s presum ption. The other specimen, indeterm inable specifically, belongs in the genus Am m odiscus sp.
Am m odiscus charoides P a rk e r et Jones
Grzybowski J., 1896, pp. 280—281, PL VIII, figs. 39—43 (slide no.
28a, b, c); recte: G lom ospira charoides (J. et P.) Pl. II, fig. 6a, b, 8
“A. charoides. Brady. Chall. rap. S. 334. Tab. X X X V III. Fig. 10—16. Tro ch a m m ina prqteus. cz. Karrer. Grzyb. Sb. d. A. W. Bd. 52. S. 494. T. I. F. 4.”
According to the opinion of R. Noth (1951) and W. Griin et al. (1964), the specimens found in the collection correspond to the species G. cha
roides (J. et P.). Besides, th e re are two specimens corresponding (Pl.
VIII, fig. 41) to the* v ariety G. charoides var. corona C ushm an et Jarvis.
Am m odiscus gordialis Jones et P a rk e r
Grzybowski J., 1896, p. 281, Pl. VIII, Figs. 44, 45 (slide no. 29).
recte: G lom osp ira gordialis (J. et P.).
Pl. II, fig. 7a, b
J. G rzybowski’s description and figure conform closely w ith the preserved specimens.
A m m odiscus Schoneanus Siddal
Grzybowski J., 1896, p. 281, Pl. VIII, fig. 46 (slide no. 30); recte:
Plectina sp.
Pl. II, fig. 9a, b
UA. Schoneanus. Brady. Chall. rap. S. 335. Tab. X X X V III. Fig. 17—19.”
“T est fin ely arenaceous w ith siliceous cem ent, surface slig h tly rough in the form of a tube coiled in a close spiral. In a fragm ent 0.8 m m in length, 5 coils, no in itial portion. V ery rare in green clays."
— 174 —
One specim en preserved in th e collection co n sists. of biserial series of cham bers arranged spirally along th e te st axis, distinct in tran sm itted light. Due to the lack of th e early portion and poorly p re served a p e rtu ra l face, this fragm ent can only be assigned to th e genus P lectina sp. I t should, therefore, be excluded from th e synonym s of T u r r it e le lla schoneana (Siddal), as suggested by V. Pokorny (1953) and F. Huss (1966).
Am m odiscus fa lla x Rzehak (Cat. E. M.) Grzybowski J., 1896, p. 281, Pl. VIII, fig. 47 (slide no. 31);
recte: ? Trocham m inoides sp.
Pl. II, fig. 10
The specim en preserved in the collection corresponds to J. G rzy
bowski’s description and figure. In tran sm itted light, globular cham bers connected by short, th in channels are visible.
A gatham m ina dubia n. sp. (Cat. E. M.) Grzybowski J., 1896, p. 282, Pl. VIII, fig. 49 (slide no. 34);
recte: R zehakina epigona (Rzehak)
Pl. II, fig. 12
The preserved specim en corresponds to G rzybow ski’s figure, y et it belongs to th e species Rzehakina epigona (Rzehak).
T roch am m in a subglobulosa n. sp. (Cat. E. M.) Grzybowski J., 1896, p. 282, Pl. VIII, fig. 50 (slide no. 35);
recte: H aplophragm oides subglobulosus (Grzybowski).
Pl. II, fig. 11
There is one small specimen in the collection which, considering its stru ctu re , should be placed in the genus H aplophragm oides. The specimen corresponds p a rtly to J. G rzybow ski’s figure, only the aper
tu ra l cham ber in the figure is too elongate and extending backward, which gives it an evolute aspect. Test subglobular. In the last-form ed whorl, 6 cham bers increasing gradually in size, separated by slightly depressed sutures. The last-form ed cham ber the largest, ap e rtu re slit- -like at th e base of the chamber. W alls of fine angular light quartz grains. S urface rough. D iam eter 0.25 mm.
