Belemnites of the genus Actinocamax Miller, 1823, from the Cenomanian of Poland

Vol. 22, No. 2

RYSZARD IMAIROI1NOIWSKI

acta geologica polonica

Waruawa 1972

Belemnites of the genus Actinocamax

Miller, ^1823, from ^the Cenomanian of Poland

ABSTRACT: The Ibelemni.tes of the genus Actin.ocamax Miller, lB23, and repre- senting iIts iSUbgenUlS Praeactinocamax Najddn, 1IJ64, recorded :fuoom <the Upper CenorIJ.aIni8!1l congl(llXl.ell"ates from G1a.naw, PollshJura Chai.n, are de6cr~bed. Two su'bspecles, PTaeactinocamax pZenu8 plenus (Blainville, 1827) and ,P. prim us p1'imU8 (Al"khangelslky. '1Qm) /Were identilfied. The s1lr:altigraphic \l"aIIlgas of ·these lbelemnites

'and :some problems 'COnnected 'With the Acti'lllOOSlll!tX plem.tS IZone arediscllSSed.

Borings of the ioh!Iro8pecies Dendrina anomala Magdefa:ollll1, 11937, and Calcidelet1'ix breviramosa Magdetkau, 1937, found on the surfa<:e

at.

,the' gu8ll."ds are :interpreted as formed postmortally, when the gu8ll"ds were par,tly 'buried 'in the sea bottom.

'JIN'I1RICIDUCTOOIN

The present paper deals with paJeontologic--stratigraphic problems concerning ; belemnites belonging to the su:bgenus

Praeactinoca~

Najdin, 1964, of the -gen'US Actinocamax Miller, 1823, recorded from the Upper Cenomanian conglomerates at Glan6w (Fig. 1), during studies on the transgressive Cretaceous deposits , in ,the' Polish Jura Ohain. The preservation of the rostra collected, which was effected both by processes active in sedimentary environment and diagenetic processes, was also taken into consideration.

The

surfac~s

of the rostra are partly corroded -by 'Pits of quartz grains and fine pebbles, like other skeletal remnants occurring

in

Ceno- manian sandstones at Korzkiew near Cracow (cf. Radwailski 1965, p. 187).

Besides pits,some borings belonging to the ichnospecies Dendrina

anomala Magdefrau, 1937 (cf. PI. 1, Fig. 2) and Calcideletrix breviram08a

Magdefrau, 1937 (cf. PI. 2, Fig. 2) were noted.

It

is of interest here, that

these 'borings ' are limited to the ventral side of one specimen (PI. 1, Fig.

la)

248

SOSNOWI£C

~

BYSZABD MABCINOWSKI

.p

-sr.

Wolbrom

C"> Clomfw

~.

y

Korzldew

.

_CRACOW

.

"%-

Fig. 1

Location map showing some Cenoma- nian exposures (dark circled) in the

Polish Jura Chain

and to the dorsal and apical part of the rostrum of the other specimen (PI. 2, Fig. Ib-c). Such distribution, as well as pelecypod encrustations (PI. .1, Fig. 3) indicate that for a long time

the

guards were only partly buried in the sediment after the animal's death

(cf.

Fig. 2) .. This made

Fig. 2

Taphonomic situation presenting the investigated guards when lying, partly buried, in tpe bo.ttom and parlly exposed

to

:the aclivity of borl.ng orga·nisms (f<Urther eltP!a-

nation in the text)

ACTINOCAMAX FROM THl!: CENOMANIAN OF POLAND 249-

possible the boring activity of rather unknown organisms over the part of the rostrum stretching out of the deposit.

In

the view of the above facts, the opinion of MagdefI'6u

(1937)

and Pugaczewska

(1965)

that.

borings

of

the ichnogenus Dendrina Magdefrau,

19·37,

'Were made by parasites during the life of the ibelemnite, seems strongly questionable

(cf.

also Najdin

1969,

Radwanski

1972).

The discussed lborings Dendrina.

anomala Magde:lirau and CaZcideletTix breviramosa Magdefrau were not.

hitherto known from deposits older

than

Senonian

(cf.

Magdefrau

1937,.

Hantzschel

1962,

Pugaczewska

1965);

the latter ichnogenus supposedly·

may also occur in the Cenomanian

of

England (Kennedy

19·70).

AcknowZedgements. The Il'IlthOlr !Sincerely :thanks iDocent A. Radwanski foc critical reading of the maarUSClriipt and helpful rema'l'lkis OOllcern4ng ithe taphionomy o-f tthe presented :belemnites.

SYSTEMATIC DESCHllPl'ION

In this chapter the taxonomy used by Jelet7Jky

(1948)

and Najdin.

(1964)

is

applied.

The biometrical data of the specimens are abbreviated as follows

(cf.

Fig.

3A, B):

S - f.O.tal length of rostrum,

LL - maximal width of I'06trum, measuxed betwee!l1 ISiIdes,

DV - maximal width· of rostrum, messuxoo from dor.sa!l to ven1lra!l 'Side,

ZZ - rostrum wildtlh measured between sides cIo.se

·to pseudoalveola,

dv - 1r000000um width me8SllJlrOO from dorsal to ventral IS'ide clQse ·to pseudoalvec>la,

LL _ index· of dorso-ventral flatteness of rostrum, DV measUTed in dts widest section,

~

^{- index} of· domo-ventr.a'l flatteneas of rostrum, dv measured close to pseudoaolveola,

LL _ index of difiell.'ence i!l1 TOStrum thickness,

Zl .

2 -

.index of :r'OS'brum elQll!gati()n.

LL

Fig. 3

A

11

s

Biometric measurements of the investigated belemnite guards A dorsal or ventral view, B side vie\,?"

8

dv

250 RYSZARD MARCINOWSKI

Family

Belemnitellidae

Pavlov, 1914 Genus ACTINOCAMAX Miller, 1823 Subgenus PRAEACTINOCAMAX Najdin, 1965 Praeaetinoeamaxplenus plenus (Blainville, 1827)

(PI. 1,

Fig. la-le)

1853. Belemrittelta plena, Blainv.; Shar:pe, pp. 9-10, Pt I, Figs 12-18.

18'12. Belemnttes lanceolatus, Sow.; Frit!lJch, p. 18, pI. 11, Fig. 6 (non Fig. '1).

1876. Actinocamax plenus, Blainv.; Schll1ter, pp. 1811-188, pt 52, F4gs 111-19.

11899. Acttnocamax plenus Blainv.; semenow, p. 93, PI. 2, Fig, 2.

1964. Praeacttnocamax plenus plenus (Blainville), 1827;· Najdin, pp. 43-46, Pi. 1, Figs 1-2;

PI. 2, Figs 1-2.

1965. Acttnocamax plenus (Blainville, 182'1); Schmid, pp. 1118-519, PI. 53, Figs 2--3.

Material. - One, nearly complete rostlrum and one fragment.

Biometr'Y (all lialear measurements in mm):

Specimen

1 76.4110.91 9.9 /1.10

_10.S

_9.6

_1.09 ^{I 7.8}

I

^{8.7 1}

^0.90

^11.40

¹

^7.

⁰¹

fligured in PI. 1, Fig. 1 (partly figured la!. PI. I, Fig. 8)

Description. - 1RcJIsrbr.tm;l cylindlrical in ven1l1'8!1 view, atta.ining maximal 1hdcknl!iSS one-thwd of the len$h ftlom the 8pi<:8!1 -end, .na:rrowi.ng from "here tOlWards the front (i.e. psoeudoalveo1:aor pa:r:t). !Ln laterai view, rOlSlbrum is wea,k1y asymmetrical,

som~t lanceolate. Dorsai lImIIr!gLn stTa:ight; apicaal end shifted somewhat toWi8X·d.9 the domal ISlide. Ventral !Side dl1aJt, whereas d<lll.'Sal S"lIiglhtly coJ;rVeX -c:1ase to pseudo- alveol-a due to latemiJ. flatteness !of Isu'balveolax par:t of the rostrum. Anterior aJIld median, dorsa!! par.ts of the 1l'0IStrum m·namented 'With lndi;s.timd, long:!twclinal striae.

Pseudoa.1veoiLa slUIbt'riangula!r .in Cl"OISI&-ISection and developed in f'arm of low cone with :shallow, and ·very small cavity near the top. Surface of pseudoalveola covered with

!Tadial ldepreslS1ions and eleVIB.tioo.s. lPseudoa[~eo1a distincb1y sepail'"ated fram the remaining parts ^of the ros1a'UiIn; ipSeudoalveolaT maD.'!gi.n parliOO!liaa."ly shaJrp on the dorsai side and descend& hell'e a few mUimeters' posteri()j['ly,. ^[aICIk of ven1lral. fissure.

Doroo-IateT.a[ fu'rr.aws fWii.de, flat ,a.nd very .indiBtlnat; conotialue posteriorly over one- -third of the rostrwn length.

Affinities. - NajdLn (1'164) estaobli:Slhed thiree Bubgenem of the gen'lllS Actino- .camax Mineit', 1823, viz. Dominative subgenus Actinocamax MUler, 1<823, and two

new 8ubgenera, Prueactinocamax lNajd:iJn ~nd ,puractinocamax Najdin, d:Lffermg in a :number 'of feaibuJres 8iDd lSItra;tiglraphic lI."a'Il!gelS. J!"·orms belonging to the su'bgenus .Praeactinocamax Najdinare characterized by a significant individual variability due to wide geographic ldistribution aJlld" perhaps, sexua'l dimolrphiml o(!Najdin 1004). The su:bgenus Pro.eUcti'liocllma.:r Najdin IDncludes two species, P. pZenus (Blainvi11e, 1827) and P. primus (AlrIkhangelsky, 1:9!12). The ISpecies ,po plenus (BLaiinv.i.iIJ.e) d4.ffer>s from P. primus (Artkhalll.ge1sky) in .genell'a1ily largEit' IS'ize and more [anceo.late shape of the .rostrum (JeletZky :19418, Birlkelund 1957), smaLler index of ll"OIStrum elongation

(~

equal 4J5~7, rarely mOIre, .in P. plenus in oomparlson "be 7-012.5 in P. primus), and aaTger 8!Pi-calJ. aJIlIgle l(Najdin ,1:964). MO'reover, the stratigraphic range is different, . because P. pri'I7IIUS (A~khangeilsky) is Iconfined to the iMiddle Cenomani"an a,nd lowet"

parts of the Upper Cenomani'an, whereaJS' P. pZenus, deri~ed firom the .fonnea-, is

ACTINOCAMAX FROM THE CENOMANIAN OF POLAND 251

known iTom ,the 'Upper Cenomaniaal and Lower TUJronlan (Je:letzky 1'948, Bidtelund 195'7, Najm..n 111984, Cbr.isibe'nsen l!9'ro) 1.

oRemaTklB. - The specimens lStudied cOn'espond to descriptions and figUl'es given 'by Fri1lsch lr19~, Pl. 1<1, Fig. 6), ISchliiter (1.8'f6, ·PI. 52, Fig. 111), a·nd Na'jdin

(l~ ,Pl, I, ~ 1--<2), ihenc:e(cf. ISYlDOOymy) their affiliation wiflh the sub;pecies Pra.eactinoca.max 1XentloS Plenu8 «BlainvLUe) .is confirmed.

Theitr

dlndex of rosbnun elong-ation !is high, thus ,they !represents fmms with Slender :r<l81:rum within this sUlbSlpecial.

Moreover; farms fil~'l'ed by SC'hmid -(1965, Pl. 53,Flgs 2---G), 'OOIlT'espondJiing to the authar'll! spec.ilmens in shape a'Dd silze' of the rostrum, ere als() induded i.n the

<synonymy. The author reexaml'ned 1lhe '9pecimen identified by CieSlitJ.skoi (1959, IP.I.. 3.

Fig. 2) IllS Acotinocamax plenus '(Bla;iJnv;LHe), bo\lIsed 1n the iMUS'e'Um oftlhe Palish Geo1ogkall Sw.-vey, 'but the specimen <is uIlISufficiently ,pres€ll'Vedfor 8ubspecific determination. The same may be said in the case of Actinocamax ple1L1L8 (I1UainvHle) previously cited' from ZIlileSJice by the present author (Marcinow.soki 100'0).

Occurrence. - PTaeactin:ocamax plenus plenU8 (!BJadnville) was found in the Upper Cenomania·n con.gI-om€lt'atas at Qlacow, '1 mete'!' 'bellOWtbe OCCU'IU'enee .site

()f Inoceramus labiattloS Schlotheim (locality no. l00c, set .2c) J. This atib&pecioes hall!

not hither,to ibeen reported from 'Poland.

'Fanne

g.enerally identified BEl Actinocamax plentloS (Blainville) 'Were cited iIn PnllliIlid from the Creta:ceoU6 depDlSi1:s of the lJWOwek lBasin '(iLawenbe:nger MuLde) .in Lower SioJ.ema .(WUoliger .1881, Scupin 119112-1'91'3., H£intzschel ::L~I33)', from the Upper Cenoma'Ilian and lowermost Turonian Qf the noNhern maxgin of :the Holy Cross 1M<Is {CieSliii'ski 1959, 1965; Cieslitiskli & Poia- ryeki '11970), from the Upper Ce:Il'OmaniBlll of iBuneIlli.n 8['ea, westenl part 'Of the

L6dz synclinorium (Cielilinski 1958) and from the Upper Cenomanian at PorE:ba Dzierzna and Slawni6w (Sujkowski 1929), and Zalesice in the Polish Jura Chain (Marcinowski 1970) ".

PTaea.ctinoca:ma:r: plenus plenus (Blalinv;i-lle) is aJso uown from the Uippelt"

Ceoomanian 1Imod 'LoWer ~oniBlll of Englam.d ~h9!IlPe t11853), ~y and Bohemia . (F11'1i:lsch

1m,

Schl.iiter ,118'716, Sclmrid .1965) ·and :K.azak!hlstan (Semenow 1399, Najdi:zi 196,1,). It ehO'Uild 'be lIlIoted thi8Jt.alfuQll'gh 1llrls s'Ulbspecies is !km:oVWl bath from Western EU!l'OIpe and Po!I.8ind ,and fram lK~aildlIs1;a·n, it 'h&s not been recorded !Tom the iEUl'n- pean pan of ·the Soviet Uni'On (cf. Najdi,n) 3.

1 Naj'din (1'9E14) defi.ned the ralIlge

of

P. p?'imus (Arkhangelsky) as uppelt' part of the [;owelt' Cenoana!lloian

to

'lowe'l' part of the UPPe'l' Cooomanian, but he applied bipartite SUibdiV'1sion 'Of rthe CetlO1llandan. Hence, it more or lea; corre.spands to the stratigre,phi<: l'lliDge given by .B.irkehmd {11l'57) and CbristerJsren {l970), a:pPiY'1ng

triparti<te is<ubdl:vision. . .

. I The alumber 'Of locaUties and lithologic members are takenforom the .authoc'lS paper (in prepall'ation) on the ,1!ramlgr$9l.ve CretaceolllS depos~ of the western maTgUn of the !Miech6w 8yniClinorium, fr·om CZE:stachowa

to

G'lanow (.Pn1ish Jura Chalin).

I Of these authors it was only Sc'Ilpin who f1gured (Sc1JPlin UIl1~Hn3, pp.

92-93, Text-fig.

8,

a broIken and poor'ly preserved ~u&rd -of Actinocama:r ~us

(B'tainviMe) ·from rthe Upper Cen.omaman (!plenus-MeU'geln) at ,J;agla:r:z «!HiIrseberg) near LWOwek; its eubspecific detexm.imation is ho:wevea.- imposBiobl.e.

4 Schliite.r ,(11871,6, p. ,1.87) aft~ ,Pusch ·(1837) ;s1:ated that this. spooies .iJs knc;>wn in Fo'land under the .nmne of Belemnites lanceolatus Sow.. w'h·lICh evidently :liS a mistake, lbecause iPusch (1837, p. 1'62) -cited BeZemnites lanceolatus? Sow~ Nom :the Dogger ·of ,the Cz~a area and therefore ;these forms s·U1'ely oCoonot belong to

the genus Actinoca.max Miller. .

, The forttl6 .reo<lll'lded :fIrom the Upper Cenomanian and Lower Turoman of the EUTqlean part of the Soviet U.nion bel-ong to different wbepecles {see below)

252 BYSZABD MABCINOWSKI

Praeaetinoeamax primus primus

(Arkhangelsky, 1912) (PI. 2, Fig.

la-le)

1912. .Acttnocama:r primus Arkh.; Arkhangelsky, pp. n8--581, Pl. 10, Figs 1-2; 4-5 (non Fig. 3).

1948. Acttnocama:r prtmuB Arkhangelsky: J'eletzky, p. 340, Text-tig. 1.

1957. Actinocama:r prtmuB primUll Arkhangelsky, 1912; Birke1und, pp. 11--12, PI. I, Figs 1-2, 4- 1964. Praeacttnocamax primus primua (Arkbana:elsky), 1912; Na:ldin, pp. ~, PI. I, Fig. ':

Pt Z, Fip 3-4, 'l.

11970. Actinocama:r primuB Arkbangellky, lS12; Christensen, pp. '16-'13, Text-fig. 2.

I

I

Material. - One, nEarly comple.te r·ostrum ~pa.rtly de8'troyedby dia·g:n:t:c pits).

Btometry (all 1iIllear measurements in mm):

speclJnen

i-·

fJlUred. ID PI. 2, Fig. 1

I ^SiLL I ^DV I ; ^{Ill! d. I-;-I~} I ^~

^t

---...:./-7.2-7-'-S.7-:-'-7.7--'-1-1.13--!.-1 6.,-, 6.9./ 0.9411.34/s.;7

Description. - ·RoW:um elongated, with index of elongation relatively high

( ^~

LL

^{= 8.27),}

s-pt.ndle .... li:ke in venok.al view, attaitning .maximal . width at one-third of the length from the apiiCaJ. end, gll.'sd·uaUy .naaowing towa~dJs ,the anterior (i.e . .pse- udoalveolau:-part). Vell'ttad 6'kie flat. !Rostrum somewhat lanceolate

in

lateral view and dorso-ventraJJ.y flattened. Dona[ side $llghtly ·f1albtaned in the Widest section, becoming eonvex in its UPPE'l' part, parti'CUlB!l'ly close to the pseudoailveod&. Where sculpture is presened, fill1e, longitudinal striae ·may 'be o'ooerved on both dOl'Sld and ventral side. Pseudoalveola in form of low cone with small depression on the top, rounded, otrapemoidaJl in cross-seo1;ion; longer axis of the :Iir.a.pewid marks dmso- -ventral di'rection, Surface .of ll6eudoalveola 'COvered with radlal depn!!S5ions and elevations, aDd concentrical rings (somewhat oblite!-ated in the investtgated speci- men). ~erud<>eJlveola lSeparated kom the .rest of Il'"ostrum wi.1lh distinct mB!l'gin.

Ventral fissure very weakly developed, in the from of short incision of pseudoal- veolaa." mll.1".gin, pas:sing opClstenody in fla·t depression, about 5 mm long. Darso-lateral furrows well develoPEd, continue up ·to haUl of the trOStrum lEngth, sligh:tiy poote- r-iorly bending towa,rds the ventil'aJ. side.

Affinities. - PTaeactinocamax primus primus (Arkhangelsky, 1.912) is S'lmiIar to the 9UbSlpeciElB P. primus elo1l.gatus (Arlthangelsky, 1912) differing in smaHer

( S .

index 'af rostrum elongation LL) and hlghelJ.' aopi-c8!l. BIIl8'le (IA1"khoogelSlky !1.91.::2, Jeletmky 19418, iBlrkelund 1957, Najdin 1004). The subspecies. P. primu.s curtus, esta- blished by Naj.d.in (1964<), differs 1com :two a'oove subspecies in commonly mOT'ter rostrum (Najdin 1984, p . .sl).

-Remarks. - The autho!I"'s specimens aa:e most comp8ll'able

to

foa.'m& figured by Bir'k:elund (112157, Pl:. 1, Fig: 1) and Najdin (,1964, P1.2, Fig. 3):.

A fragment of a very slender rostrom, och8lraclerized by a very low apical angle. found by the a'Utbo.r, seems ,to be related to P. primus 'elongatus (Ar'khangel- sky) but :it is too inOOmplete for determinat.ion with certalini-ty.

B~kelU!lld (lQY1) suggested that all J;pecimens fig·uTed ,by. ArkhangeIslky (19'12) as Actinocannnx primus Arkh. (A!I'"kbangelsky 1¹91:2, pp. 57'8--0581, P!l. 1.0, Figs 1-5) and ditfelJ.' frQUl ,typical P. plenus pl.enus .(iBlainvill.e) in pseud.oa1veola dndi&thDctIy separated from the rest of rO&tIrum, a:nod eommonJ.y in a nm.rower iSuba!lVerolar part and a slightly cancave anterw part of the rostrum (Najdin 1964).

ACTINOCAMAX FROM THE CENOMANIAN OF POLAND

belong to the subspecies P,raeactinocamax primus primus (Arldlange1sky). Howe- ver, aCOOIding to ·Najod:in {1964) , Atklha.ngeLsky's 'SlPEcimen figured in bile Pl. 10, Fig. 3 shou:ld be imc1cuded ,to the :subspecies P. primU8 elongatus (.A.1"khaDlgelsdty);

the 1atte'l' vieW'POilllt -iIs 'accepted by the present author.

ChTistensen (lJ970) givES no reason fori.dentify~ng his specimetIB

to

spedfre l'aIlIk only; it follows from the sYlllooymy and description of the material that these speciroeilllS moOSt ,pr-obably ISOOWdbe -included to the ,subspecies P. primus primus

(.Arlkhang~"'ky).

Occurrence. - Praeactinocamax primus primus (Arkhangelsky) was rec-or- deod from the s-ame -Layer of !the Uppe!I' Cenomanian -congllOotn-erates at Glanaw as P. plenu8 plenus (localil1y 00. IlOBe,

:set

-2c). The species and 6uoopECi-es undEl" dis- cussioo i'8 cited for the fllr.st :time fr·om Poland.

Praeactinocamax primus primus (A'l'k;hangelsky) WaIS -hitherto known from the Middle 8JIJ.d !lower part of the Upper Cenoman~an of .Demnall'k o(1BiTkelund 1957), Scania (Christensen 1970) and number of localities in the Eu,ropean part of the Soviet Uni{)'Il and lKazakhstan {A'l'khangelSky 119112; Najdi.n 1'96D. 1964).

STRATIGRAPHIC SETTING OF DEPOSITS YIELDING .PRAEACTINOCAMAX P4ENUS (BLAINVILLE)

Stratigraphic setting of deposits with Praeactinocamax plenus -(Bla- inville) and hence the vertical range of this species is still controversial.

Some authors include all deposits yielding this species, i.e. the Actinoca- max plenus Zone, to the Upper Cenomanian (Barrois 1675, Petrascheck 1905, Hanoock 196!:l), whereas others include these deposits to the Lower Turonian (Hebert 1874, Spath 19-26, Hantzschel 1933, Jefferies 1963). In turn, the

third

group of authors, particularly those studying the Cretace- ous of Bohemia and Saxony, hold that P. plenus (Blainville) occurs in transitional beds betWeen the Cenomanian and Turonian (Pietsch, fide Schmid 1965) and regard these beds as the separate ActiIwcamax plenus Zone

(cf.

Tr5ger 1967-19'69), as do some English .authors

(cf.

Kennedy 1969) .

. Moreover, correlation of deposits

of

the Actinocamax plenus Zone raises certain doubts. Troger (1969) states tliat Metoicoceras geslinianum (d'Orbigny) has been occasionally recorded.in Saxony and Bohemia from the Upper part of this zone; hence, that upper part would 'correlate with the lower part of this zone

in

south-western and south-eastern England (cf. Hancook 1969; Table 1; and Kennedy 1969, Table 1).

In Najdin's opinion (1964), more accurate evaluation of the strati- graphic rang.e of P. plenus (Blainville) will be posSible after paleoritologic elaboration

of.

this index species, which comprises several subspecies dif- fering in stratigraphic ranges. For example (cf. Najdin 1964), Praeactino- camax plenus plenus (BlainviUe), P. plenus longus Najdin, P. plenus acu-

tU3

Najdin

^OCC'UT

in the Upper Cenomanian and Lower Turonian, whereas

the occurrence of P. plenus triangulu8 Najdin, P. plenus.CTas8u8 Najdin

254 RYSZARD MARCINOWSKI

and P. plenus contractus Najdin is limited to deposits with Mammites nodosoides (Schlotheim) and Inoceramus labiatus Schlotheim of the L0- wer Turonian.

"At Glan6w, P. plenus plenus (Blainville) was recorded

in deposits

yieldin'g Schloenbachia d. lymensis Spath, "the species Ilmown from the Upper Cenomanian only (cf.

Spa~h

1926, Wright & Wright 1951, Ciesoo- ski 1959, Kennedy 1969), and P. primus primus (Arkhangelsky). Co-oc- cwrreoce of P. pZenus plenus (Blainville) and P. primus primus (Arkhan- gelsky) indica'tes that these deposits 'beiong

to

the lower part of the Acti- nocamax tplenus Zone, since the fOMler subspecies is known eJrelusively from the Upper Cenomanian and Lower Turonian, and the latter ':from the Middle

"

and lower part

of

the Upper Cenomanian

(cf.

Jeletzky 1948, Bir- kelundl957, Najdin 1964, Christensen 1970).

At Glan6w, the upper part of the lithological member yielding the belemnites under discussion. exhibits distinct traces of scouring 'and is overlied by laminated limestones. Therefore

it

may be presumed that depOsition was impeded for a period of time in "the sedimentary basin in the Glan6w area, which resulted in a partial removal of the deposits, and this

~ess

was followed by a rapid change of lithofacies

in

the latest Cenomanian.

Institute

of

GeoZog'U

"

Of

the Wa1'8'I1W U'llilveTsoit'J/

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~~~ .

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R. MARQIlNOWSK:E

BELEMNITY Z RODZAJU ACTINOCAMAX MILLER. 18!3, Z CENOMANU POLSKI

(Streszczenie)

W pracy op_no belemnity {pl. 1, fig. la-le; pI.. 2, fig. la-le) Z Iodzaju Aetinocamax Miller. 1823, .n.aleZ~ce do pod'l"Odzaju Praeactinoeamax Najdin, 1964, .a mianowicie Praeaeti1WCa.max plenus plenus (Blal1n,,'ille, 1827) i P. pTimus primus (Arkhangelsky, 1912), kt6re znaiezione zostaly w ' g6mocenomaflskich zlepieflcach w Glanowie ^I(fig. 1). ()1I:nibwi:ono .za.si~gi IStratYgr.afl'c7llle -tY'ch belemnit6w -or,az ^Z9.- gadnienia %Willzane z pozycjll str·atygraficm~ poziomu Actinocamax plrnUlS. Anali- zujl\C wyst~pujllce lIla powierzcbni badanych rostraw Mllzenia (pt 1, fig. 2; pl.2, fig.

2) lIlaleUl'Ce

do

ichnogatu.nk6w Dendrina. anomaZa Mllgdmau, 1-937, ·oraz Calciaeletrix oreviramosa lMilgdef!reu, 1937, 7lWr600no uwagf;l, iZ utworzoOne one zost~y po Smi-srci belemnitaw, wOwezoas edy 'I'()S1lrum "l!poczywalo ·na dnle, CZf;l9ciowo pogrllzone w ·osa- dzie (f1g. 2).

Instytut Geologii,Podstawowej Uniwersytetu Wa'rszawskiego Warszawa 22, Al. Zwirki i Wigury 93

Warsztiwa,

w

listopaazie 1911 T.

ACTA GEOLOGICA POLONICA, VOL. 22 R. MARCINOWSKI, PLo 1

10 lb le

la-c - Praeactinocamax pZenus plenus (Blainville); Glan6w, Upper Cenomanir.:l - a v ntral view, b side view, c dorsal view; X 1.5.

2 - Boring Dendrina anomaZa Magdef.rau cin the gual'd of Praeactinocamax pZenus pZenus (Blai.nvme) pn entec1 in F'l.g. 1 of this plate, X 10.

3 - EpizOic peleeypod, Dimyodon nilssoni (v. Hagenow), on the other guard of Praeactinoeamax pZenus plenus (Blainville)' Glan6w Upper Cenomanian, X 3,

All photos taken by B. Drozd, M. Se.

ACTA GEOLOGICA POLO ICA, VOL. 22 R. MARCINOWSKI, PLo 2

10 lb le

la--c - Praeactinocamax primus primus (Arkhangelsky); Glan6w, Uppe·r Ceno- manian - a ventral view, b side view, c dorsal view (the guard partly corro- ded by diagenetic pits); X 1.5.

2 - Borings Calcideletrix breviramosa Magdefrau in the apic~l part of the guard of Praeactinocamax primus primus (Arkhangelsky) presented in Fig. 1 of this plate, X 5.

All photos taken by B. Drozd, M. Se.