R O C Z N I K P O L S K I E G O T O W A R Z Y S T W A G E O L O G I C Z N E G O A N N A L E S D E L A S O C I E T E G E O L O G I Q U E D E P O L O G N E Tom (Volume) XXXVII — 1967 Zeszyt (Fascicule) 2 K raków 1907
EW A ŁU C Z K O W SK A
KILKA NOWYCH GATUNKÓW OTWORNIC Z MIOCENU POLSKI
(Tabl. VIII, I X i 7 fig.)
Some new species of Foraminifera from the Miocene of Poland
(PI. VIII, I X a n d 7 Figs)
STRESZC ZEN IE
Autorka opisała 5 nowych gatunków otw om ic znalezionych w osadach miocenu na obszarze m iędzy Tarnobrzegiem a Chmielnikiem na południe od Gór Świętokrzyskich i w ystępujących również na innych obszarach w przybrzeżnych osadach miocenu Polski południowej. Gatunki te b yły cytow ane w pracach autorki (1964, 1966) jako noimina nuda i nie zostały dotychczas dokładnie opisane. Ich położenie stratygraficzne jest podane na załączonej tabelce (fig. 1). Dwa gatuinki można uważać za charakterys
tyczne dla miocenu Polski: Anomalinoides dividens dla dolnego sarmatu w facji ilastej i Textulariella lithothamnica dla dolnego tortami w facji litotam niowej. Użyta w cytowanych pracach autorki nazwa Textulariella tortonica została obecnie zmieniona na w łaściwszą T. lithothamnica.
A b s t r a c t . F iv e n e w sp ecies of F oram in ifera, fou n d in th e M io c e n e se d im e n ts so u th of the H o ly -C ro ss M ou n tain s are described, i.e. T e x t u l a r i e l l a lit h o th a m n ic a , B u l i m i n a angusta, B u li m i n a m ic ra , G l a b r a t e l l a p l a n a and A n o m a l i n o i d e s d i v i d e n s . T h ey w a s cited in au th or’s papers ( L u c z k o w s k a 1964, 1966) as n o m in a nuda.
IN T R O D U C T IO N
F ive species of Foraminifera are described, found in the Miocene sedim ents of Tarnobrzeg and Chmielnik region, south of the Holy-Cross M ountains and distributed in other areas of near-ishore sedim ents of Miocene of Poland. These species are cited in author’s papers (1964, 1966) as nomina nuda and hitherto w ere not in detail described.
The stratigraphical position of the described species is shown in Table 1.
Two of them can be considered as index fossils in the Polish Miocene:
Anomalinoides dividens in the low er Sarmatian clays and Textulariella lithothamnica in the lower Tortoniam Lithothamnium-limestones. The nam e Textulariella tortonica used in the papers quoted above is changed to T. lithothamnica as the latter nam e is considered to have more signi
ficance for the species.
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T a b l e 1
A g e Zone Sub zon e S p ecies
L o w er S arm atian
E l p h i d i u m h a u e r i n u m Q u in q u elo c u lin a s a r m a t i c a
G l a b r a t e l l a p la n a B u l i m i n a m i c r a Q u in q u elo c u lin a
k a r r e r i o v a t a A n o m a l in o id e s d i v i d e n s
A n o m a l i n o i d e s d i v i d e n s
U pper T ortonian
C ib icid e s c r a s s i s e p t a t u s
N e o b u l i m i n a longa S p ir ia lis
L ow er T ortonian
U v i g e r i n a co sta i
C a n d o r b u li n a u n i v e r s a A m p h i s t e g i n a les son ii
T e x t u l a r i e l l a l it h o th a m n ic a B u l i m i n a a n g u s ta
SYSTEMATIC DESCRIPTIONS
S u p er fa m ily Li'tuolacea d e B l a i n v i l l e , 1825 F a m ily P a v o n itin id a e L o e b l i c h & T a p p a n , 1961 S u b fa m ily P a v o n itin in a e L o e b l i c h & T a p p a n , 1961 T e x t u l a r i e l l a C u s h m a n , 1927
Textulariella lithothamnica n, sp.
(PI. V III, fig. 1— 5, PI. IX , fig. 7 t e x t - fig s 1, 2)
H o l o t y p u s : PI. VIII, fig. 1, N o F 601.
L o c u s t y p i c u s : Grzybów near Staszów.
S t r a t u m t y p i c u m : Lower Tortonian, Lithotham nium -lim estones south of the Holy-Cross Mountains, subzone w ith Amphistegina lessonii.
D e r i v a t i o n o m i n i s : from the Lithothamnium-beds.
M a t e r i a l : above 50 specimens.
D i m e n s i o n s : holotypus 2,5 mim, paratypes 0,75— 3,00 mm.
D i a g n o s i s : Shape subcylindrical, the initial part w ith inflate cham bers increasing rapidly in size as added, above a quarter of the height of the test chambers low, flat, of the same shape, the number of internal partitions ranging from tw o in the initial part to six in the last chambers.
Wall agglutinated, w ith tw o layers.
D e s c r i p t i o n : Test large, subcylindrical, elongated, the initial end acute, the proximal end oblique. Number of chambens 8— 15 on each side.
The first w horl composed of three globular chambers undivided inside, they are visible outside as a sw elling at the acute end of the test. N ext chambers up to a quarter of the height of the test distinct, inflated, increas
ing rapidly in size as added, arranged in tw o horizontal series, the internal
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partitions marked at the surface of the test as sm all cavities on the chambers, the remaining chambers of the sam e shape, flat, low and smooth. Chambers internally subdivided by radial partitions near the periphery, the number of w hich increased from two in the initial part to six in the last chambers (Fig. 3). W all agglutinated, composed of two
F ig. 1. T c x t u l a r i e l l a lit h o th a m n ic a , a — fro n t v ie w , b — s id e v ie w ; G rzyb ow 27, d ep th 235— 238 m.
d e
Fig. 2 a— j. T e x t u l a r i e l l a lit h o th a m n ic a , d e v e lo p m e n t of in te r n a l p a rtition s (serial sectio n of one s p e c im e n ); 1. c., depth <235— 23i8 m .
layers, the inner calcareous, smooth, perforated, w ell visible in broken ispecimens, and the outer agglutinated w ith thin sand grains and much calcareous cem ent (P. II fig. 6). Within the chambers the pseudochitineous lining. Aperture as a low arch at the inner margin of final chamber.
R e m a r k s : Shape of the test resem bles Textularia recta C u s h m a n , but the characteristic w all of this species is agglutinated from thick calcareous grains. Textularia barretti P a r k e r et J o n e s differs in
— 236 -
having the aperture surrounded by the serrate border and the test com pressed in opposite direction than in other species of Textularia. The initial part of Textulariella lithothamnica resembles Textularia pseudo - rugosa L a c r o i x , but the latter differs in lacking the internal partitions within the chambers.
S u p er fa m ily B u lim in a c e a J o n e s , 1875 F a m ily B*ilim inidae J o n e s , 1875 S u b fa m ily B u lim in in a e J o n e s , 1875 B u li m i n a d ’O r b i g n y, 1826
Bulimina angusta n. sp.
(PI. VIII, fig. 6— 9, te x t - fig . 3)
H o l o t y p u s : PL VIII, fig. 6, N o F 606.
L o c u s t y p i c u s : Grabki D uże near Staszów.
S t r a t u m t y p i c u m : Lower Tortonian shales and clays, zone w ith Candorbulina universa.
D e r i v a t i o n o m i n i s : from the word angustus = narrow.
M a t e r i a l : about 50 specimens.
D i m e n s i o n s : holotypus 0,35 mm., para types 0,35— 0,55 mm.
D i a g n o s i s : Shape elongated, th e inflated chambers of initial part arranged in three series, later chambers being more elongate w ith tendence to be uncoiled. The last chamber narrow.
Fig. 3. B u l i m i n a ang usta, cham bers ou tlin e; G rabki D u że N 8, depth 189— 191 m.
D e s c r i p t i o n : Test elongated, tapering at both ends. The initial part triserial with sm all inflated chambers and depressed sutures, above the half of the height of the test chambers strongly increasing in size as added, elongate and w ith tendence to become uncoiled and nearly biserial. The last chamber elongated, narrow. Surface smooth, polished, fine perforated.
Aperture elongate, loop-shaped below the top of the last chamber.
R e m a r k s : The species remind Bulimina elongata d ’O r b i g n y fi
gured by C u s h m a n (1947, pi. 25, fig. 16a, 16b), but differs considerably from B. elongata d’O r b i g n y . J. A. C u s h m a n does not m ention the origin of the figured specimens. From B. elongata d’O r b i g n y it differs in having more inflated chambers, more elongated last few chambers and the shape of the test more fusiform, not broadest near the apertural end.
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Bulimina micra n. sp.
(PI. VIII, fig. 10— 13, t e x t - fig . 4)
H o l o t y p u s : PI. VIII, fig. 10, No F 610.
L o c u s t y p i c u s : R ytw iany near Staszow.
S t r a t u m t y p i c u m : Lower Sarmatian, K rakowiec-clays south of the Holy-Cross Mountains, zone w ith Quinqueloculina sarmatica.
D e r i v a t i o n o m i n i s : from the word micros = small.
M a t e r i a l : 30 specimens.
D i m e n s i o n s : holotypus 0,38 mm, paratypes 0,28— 0,58 mm.
D i a g n o s i s : Shape cylindrical, chambers subcircular, flat, compact, sligh tly increasing in size as added, sutures flat, in the upper part som e
w hat depressed.
Fig. 4. B u l i m i n a m i c r a , ch am b ers outline; R y tw ia n y 1, depth 66—€7 m.
D e s c r i p t i o n : Test small, elongated, cylindrical, at both ends round
ed. Chambers subcircular, not inflated, compact, gradually increasing In size as added, sutures flat, in the upper part of the test som ewhat depressed. Last chamber small, rounded, Surface smooth, polished, with fin e and dense pores. Aperture sm all, loop-shaped, below the top of the last chamber.
R e m a r k s : Large specimens resembles Bulimina gracilis C u s h m a n , but differs in having less inflate chambers and the aperture placed below th e top, rather than toward the center of the face.
S u p e r fa m ily D iscorb a cea E h r e n b e r g , 1838
F a m ily G la b r a te llid a e L o e b l i c h & T a p p a n , 1904 G l a b r a t e l l a D o r r e e n , 1948
Glabratella plana n. sp.
(PI. V III, fig. 14— 17, t e x t - fig . 5)
H o l o t y p u s : PI. VIII, fig. 14 a, b, No F 614.
L o c u s t y p i c u s : Zrecze near Chmielnik.
Fig. 5. G l a b r a t e l l a pla na , cham bers outline;
Z recze 3, depth 59,3— 59,4 m.
— 23 8 —
S t r a t u m t y p i c u m : Lower Sarmatian, K rakowiec-clays south of the Holy-Cross Mountains, zone w ith Quinqueloculina sarmatica.
D e r i v a t i o n o m i n i s : from the word planus = flat.
M a t e r i a l : above 50 specimens.
D i m e n s i o n s : holotype 0,32 mm, para types 0,20— 0,35 mm.
D i a g n o s i s : Shape lenticular, stellate, chambers of the last whorl radially elongate and tubulospiinate.
D e s c r i p t i o n : Test trochospiral, small, lenticular, central part of the spiral side slightly convex, opposite side umbilicate. Chambers of the last whorl strongly increasing in size, radially elongate and tubulospinate, making th e stellate outline of the test. The spines of previous chambers nearly horizontal, som etimes attached to the chambers w all of the last whorl and slightly visible. Number of chambers 5 in the early part, 6— 8 in the last whorl. Sutures som ewhat depressed. Surface of the spiral side smooth, polished, distinct perforate, opposite side covered w ith radial tuberculate striae, umbilical depression often with pustules. Aperture a small umbilical slit, open to the central depression and som etim es sligh tly visible.
R e m a r k s : From Glabratella imperatoria (d’O r b i g n y) it differs in having the more flat test of stellate outline, radially elongate chambers and horizontal spines, whereas G. imperatoria has a conical test, globular chambers and spines raised upwards.
According to L o e b l i c h & T a p p a n (1964) schizont forms of this genus are „generally larger and flatter, gamont more highspired”. It seem s howewer, that G, plana cannot represent the schizont form s of G. imperatoria on account of different shape of chambers and different horizontal spines.
S u p er fa m ily C assid u lin a cea d ’O r b i g n y , 1839 F a m ily A n o m a lin id a e C u s h m a n , 1927
S u b fa m ily A n o m a lin in a e C u s h m a n , 1927 A n o m a l i n o i d e s B r o t z e n, li942
Anomalinoides dividens nom. nov.
(Pl. IX f ig 1— 6, te x t - fig s 6, 7)
1884 A n o m a l i n a g r o s se r u g o s a G u m b e l , Sp.: B r a d y H. B., 1884, PI. 94, fig . 4, 5 193i3 A n o m a l i n a n. sp. aff. g r o s se r u g o s a B r a d y (non G u m b e l ) (n. sp. aff. T h a l
m a n n); T h a l m a n n H .E ., 1933, p. 252
H o l o t y p u s : PI. IX fig. 1 a— c, No F 618.
L o c u s t y p i c u s : Zrecze near Chmielnik.
S t r a t u m t y p i c u m : Lower Sarmatian, K rakowiec-clays south of the Holy-Cross Mountains, zone with Anomalinoides dividens.
D e r i v a t i o n o m i n i s : from the word divido = divide (it divides the Sarmatian microfauna from the Tortoinian one).
M a t e r i a l : above 1000 specimens.
D i m e n s i o n s : H olotype 0,46 mm, paratypes 0,21— 0,70 mm.
D i a g n o s i s : Shape circular, lobulate, periphery rounded, central part of the spiral side often depressed, chambers of the last whorl enlarging rapidly as added, few last chambers inflated.
D e s c r i p t i o n : Test low trochospiral, circular, lobulate, periphery broadly rounded, especially in the last part of the whorl. Spiral side co n v ex ,w ith the slightly depressed central part, last whorl broad w ith
--- 239 —
7— 10 inflated chambers, enlarging rapidly as added, earlier whorls small and sligh tly visible in the middle of the depressed part, often covered w ith thickened shell material, sutures oblique, arcuate. Opposite side involute, umbilicate, few last chambers strongly inflated, triangular, sutures radial. Wall thin, transparent, smooth, densely perforated, espe-
Fig. 6. C om parison of cham bers outline: 1, 2 a— c — A n o m a l i n a g r o s se r u g o s a G u m - b e 1, sp. (in B r a d y ’s w o r k )r 3, 4 a— c — A n o m a l i n o i d e s d i v i d e n s ; a — sp ira l side,
b — u m b ilic a l side, c — sid e v ie w
Fig. 7. A n o m a l i n o i d e s d i v i d e n s , va ria b ility of la st ch am b ers ou tlin e; Z recze 3, depth 72,7— 72,8 m.
cially on the spiral side, the umbilical side with the earlier part more sparsely perforated or nearly devoid of pores, granular in structure (PI. II fig. 7). Aperture peripheral, large, semicircular, with slightly thickened lip, extending onto the spiral side.
V a r i a b i l i t y : The morphological variability concerns the follow ing features: a) various size and convexity of the last chamber, b) various perforation of the few last chambers, particularly on the umbilical side, c) size of the test. There exist small specimens with nearly nonperforate um bilical side and som ewhat inflated chambers, besides the larger forms w ith dense pores on the w hole surface and strongly inflated last chamber.
A fter breaking of few last chambers appear the sparsely perforated hyaline w all of the initial part of the test. The periphery of this discovered part is more flattened sim ilarly as in small specimens' of this species. It seem s that the variability described above is connected w ith ontogeny.
— 240 —
The small flattened specimens w ith sparse perforation are juvenile forms and the formation of globular chambers and dense perforation is: cor
responding w ith the final phase of chamber formation.
R e m a r k s : Anomalinoides dividens is most approaching to the recent form described by H. B. B r a d y (1884) from the Atlantic and Pacific Ocean, nam ely Anomalina grosserugosa G i i m b e l , sp.; H. E. T h a l - m a n n (1933) in the n ew nom enclature to B r a d y ’s work stresses upon that it is quite distinct from A. grosserugosa G ii m b e 1, but he does not give the other name for this n ew species. In B r a d y ’s work the fu lly description is absent except the notice (p. 673), that „the w alls are coarsely perforated, but have few er pores on the superior than on the inferior side” and that Anomalina badenensis d ’O r b i g n y is a v ery similar variety. This notice as w ell as the illustrations given by B r a d y cor
respond to our specimens.
Anomalina badenensis d’O r b i g n y differs in having evolute arran
gem ent of chambers on the both sides, thick and nontransparent wall, more coarse perforation and small peripheral aperture.
D e p a r t m e n t of P a l a e o n t o l o g y School of M in in g a n d M e t a l l u r g y C r a c o w
W Y K A Z L IT E R A T U R Y R EFER EN CES
B r a d y H. B. (Ii834), R eport on th e sc ie n tific resu lts of th e v o y a g e of H. M. S.
„ C h a llen ger” during t h e years 1873— 76. Zool., 9. London.
C u s h m a n J. A. & P a r k e r F. L. (1947), B u lim in a and related fo r a m in ife r a l g en era.U. S. Geol. S u r v . Pr of . P a p e r 21'0-D, p. 55— 160, W ashington .
E l l i s B. & M e s s i n a A. (1940), C a talo gu e of F o ram in ifera , Sp. Publ., A m e r . Mus. Nat . Hist., N e w York.
L o e b l i c h A. R. Jr., & T a p p a n H. (1964), Trea'tise on in v e rte b r a te p a leon tolog y, P art C, P r o t i s t a 2, 1, 2, 510 pp., N e w York.
Ł u c z k o w s k a E. (1964), S tra ty g ra fia m ik ro p a leo n to lo g iczn a m io c e n u w r e jo n ie T arnobrzeg— C h m ieln ik (The m icro p a leo n to lo g ica l str a tig r a p h y of th e M io cen e in th e region of Tarnobrzeg— C h m ielnik). P A N Oddz. w K ra k o w ie, K o m . N a u k Geol., Pr. geol. 20, W arszaw a.
Ł u c z k o w s k a E. (1967), P a le o ek o lo g ia i str a ty g r a fia m ik ro p a leo n to lo g iczn a m io cen u ok olic G rzy b ow a k oło S ta szo w a (P a leo eco lo g y and m ic r o p a leo n to lo g ic a l s'tratigraphy of th e M iocen e in th e v ic in ity of G rzybów near Staszów ). A c t a geol. pol. 17, 1, W arszaw a.
T h a l m a n n H. E. (1933), N ach tra g zum N o m en cla to r zu B r a d y ’s T a fe lb a n d der F o ra m in iferen der ,,C h allenger”-E x p e d itio n . Ecl oga e geol. H elv ., Bd. 26, 2, 251—
— 255, B asel.
E X P L A N A T IO N OF P L A T E S V III—IX
P la te V III
Fig. l a , b. T e x t u l a r i e l l a l it h o th a m n ic a n. sp.; h olotyp e, a) fr o n t v ie w , b) s id e v ie w ; G rzybów 27, d ep th 235—238 m, X25.
Fig. 2— 3. T e x t u l a r i e l l a l i t h o t h a m n i c a n. sp., para'types; I.e., d ep th 235—238 m, 2 — X 24, 3 — X19.
— 241 —
F ig. 4. T e x t u l a r i e l l a l i t h o t h a m n i c a n. sp., top v ie w sh o w in g ea rly coil; G rabki D u że N 8 , depth 186— 187 m, X32.
Fig. 5. T e x t u l a r i e l l a l it h o th a m n ic a n. sp., fro n t v ie w , th e a g g lu tin a te la y e r d e stroyed, p erfora tio n of th e in n er la y e r v isib le ; 1. c., depth 186— 187 m, X25.
Fig. 6 a, b. B u li m i n a a n g u s t a n. sp., hole/type; G rabki D u że N 8, d ep th 189— 191 m, X104.
Fig. 7— 8 a, b. B u li m i n a a n g u s t a n. sp., paratyp es; I.e., dep'th 189— 191 m, X104.
Fig. 9. B u li m i n a m i c r a n. sp., holotype; R y tw ia n y 1, depth 66— 67 m , X84.
F ig. 10—il2. B u l i m i n a m i c r a n. sp., p aratypes; I.e ., depth 66— 67 m, X84.
Fig. 13 a, b. G l a b r a t e l l a p l a n a n. sp., holotyp e; a — spiral side, b — u m b ilica l side;
Z recze 3, depth 59,3— 59,4 m, X114.
Fig. 14— 15 a, b. G l a b r a t e l l a p l a n a n. sp., p aratyp es; a — spiral side, b — u m b ilic a l side; 1. c., depth 59,3— 59,4, X114.
P la te IX
Fig. 1 a—c. A n o m a l i n o i d e s d i v i d e n s nom . nov.; holotyp e; a — sp ira l side, b — u m b ilic a l sid e, c — s id e v ie w , X70.
Fig. 2—'5 a— c. A n o m a l i n o i d e s d i v i d e n s nom . nov., p aratyp es; a — sp iral side, b — u m b ilica l side, c — sid e v ie w , 4— 5 ju v e n ile form s, u m b ilic a l s id e w ith sp a r sely p erforated w a ll, X70.
Fig. 6—7. T e x t u l a r i e l l a lit h o th a m n ic a , th in sectio n s sh o w in g tw o la y er s, X40
Fig. 8— 9. An om,alin oides d i v i d e n s , thin section s, granular stru ctu re of 'test v isib le;
8 — X110, 9 — X168
A ll sp e c im e n s of A n o m a l i n o i d e s d i v i d e n s from th e b o r e -h o le Z recze 3, depth 72,7— 72,8 m.
